These are the main characters of the four An American Tail movies and the series Fievel's American Tails.
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The Mousekewitz Family
Voiced by: Phillip Glasser ("An American Tail", "Fievel Goes West", "Fievel's American Tails"), Thomas Alexander Dekker ("The Treasure of Manhattan Island", "Mystery of the Night Monster")
Fievel is the main character of the series. He's a fearless child, but his youthful curiosity often gets him into a lot of trouble, and he tends to get separated from his family on multiple occasions.
Voiced by: Amy Green ("An American Tail"), Cathy Cavadini ("Fievel Goes West", "Fievel's American Tails"), Lacey Chabert ("The Treasure of Manhattan Island", "Mystery of the Night Monster")
Tanya is Fievel's older sister. Tanya is a dreamer, much like her brother. Her personality is explored mainly in Fievel Goes West, where we learn she wants to be a famous singer and actress. Her personality varies from movie to movie, however, as does her appearance.
Voiced by: Cathy Cavadini ("Fievel Goes West", "Fievel's American Tails")Fievel's baby sister. She's never a huge part of the plot, and often is inserted just to look cute. The animators even forgot about her halfway through the first movie.
Mama and Papa Mousekewitz
Mama Voiced by: Erica Yohn; Susan Silo ("Fievel's American Tails"); Jane Singer ("Mystery of the Night Monster")
Papa Voiced by: Nehemiah Persoff; Lloyd Battista ("Fievel's American Tails")
Fievel's parents. Papa likes to tell tall tales and wholeheartedly believes that there are no cats in America at the beginning of the first film. Mama, however, is much more down to Earth and isn't afraid to tell Papa when she thinks his stories are nothing but fairy tales.
Characters Debuting in "An American Tail"
Voiced by: Dom De Luise
The only nice cat Fievel meets, and one of his best friends. He's a bit cowardly but he can be brave when he wants to. He's also a vegetarian, so he never eats mice. Fievel Goes West is where he saw the most character development.
Voiced by: Pat Musick
A streetwise orphan that Fievel befriends in the first film while imprisoned in a sweat shop. After Fievel frees everyone Tony takes it upon himself to help Fievel find his family. He's largely absent from Fievel Goes West except for some blink-and-you-miss-it cameos, but he comes back in full main character status in the direct to video sequels.
Voiced by: Cathianne Blore
Bridget is an Irish mouse-rights activist who Tony meets and falls in love with. She tries to use her ties with the mouse politicians in New York to help find Fievel's family.
Warren T. Rat
Voiced by: John P. Finnegan
A crooked rat with a stranglehold on the mice of New York. He is greedy, selling poor little children like Fievel into sweatshops in exchange for their salary, and running a protection racket on the mice offering them protection from the cats in exchange for money.
Voiced by: Will Ryan
A tiny roach who works for Warren as an accountant. He is generally dissatisfied with his job, but seems to be too afraid to quit. He emits sparks from his antennae when excited.
Voiced by: Madeline Kahn
A rich uptown mouse who is committed to ridding New York of its cat menace, and finally gets an idea on how to accomplish this from one of Fievel's fairy tales.
Voiced by: Christopher PlummerA French pigeon who encourages Fievel to seek his family when he washes up on Ellis Island.
Characters From "Fievel Goes West"
Voiced by: James Stewart
A retired law dog that Fievel idolizes before he moves out west. He then teams up with Fievel and trains Tiger to take his place.
Voiced by: Amy Irving ("Fievel Goes West"); Cynthia Ferrer ("Fievel's American Tails")
Tiger's ex girlfriend who leaves him to go west and have real adventures. Ends up as head matron to Cat R. Waul's saloon, and shows Tanya the glamor of stardom.
Cat R. Waul
An evil British cat who schemes to eat all the mice of Green River by luring them there with the promise that out west, cats are nice to mice. He's also Wicked Cultured and planned to spare Tanya from the fate of the other mice because he liked her singing voice.
Voiced by: Jon Lovitz ("Fievel Goes West"), Dan Castellaneta ("Fievel's American Tails")
Cat R. Waul's tarantula sidekick who gleefully obeys Waul's orders, and has fun doing it too.
Voiced by: Kenneth MarsOriginally just a random member of Waul's gang, he becomes a bigger threat in Fievel's American Tails.
Characters From "The Treasure of Manhattan Island"
Voiced by: Elaine Bilstad
Cholena is the daughter of the Chief of an underground tribe of Native American mice, who fled underground with the arrival of European settlers in Manhattan. Fievel befriends her and brings her up to the surface in an effort to prove that European mice aren't bad.
Voiced by: Ron Perlman
One of three corrupt owners (the other two are listed below) of a cheese factory, who turn it into a sweatshop and later stir the workers into a crusade against the Native American mice living beneath the city.
Voiced by: Tony Jay
Voiced by: Richard Karron
Voiced by: Sherman Howard
A crooked cop who is on the payroll of the three villains, doing their dirty work and enjoying every minute of it.
Voiced by: Rene Auberjonois
An archeologist from the Museum of Natural History that Tony worked for once, who deciphers the map Fievel and Tony find and accompanies their expedition for the 'treasure' under Manhattan that appears on the map.
Voiced by: John Kassir
Dr. Dithering's hired assistant, who as it turns out, was also secretly under the employment of Grasping, Toplofty and O'Bloat.
Characters From "Mystery of the Night Monster"
Voiced by: Susan Boyd
Nellie Brie is a no-nonsense, down to Earth reporter with a mind for facts. She's also one of the craftiest reporters New York's mice have ever known, and her work for the Daily Nibbler is legendary. She's constantly at odds with her editor, who believes more in selling newspapers than bringing people facts.
Voiced by: Robert Hays
He is the editor of the Daily Nibbler, a fast-talking and charismatic boss who's constantly at odds with Nellie Brie over whether her ideas for reports will sell newspapers.
Voiced by: Candi Milo
Madame Mousey (pronounced 'Moo-say') is a small french poodle, who fled her owner to cause some trouble on her own. She has a very short-fused temper, and starts her own gang of cats in the sewer.