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* BookEnds: The first Successor the party encountered was the Successor of the Ribcage, which looks like a wolf merged into the body of a woman. The Skull King is a werewolf-like creature.

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* BookEnds: The first Successor the party encountered was the Successor of the Ribcage, which looks like a wolf merged into the body of a woman. The Skull King is a werewolf-like creature. Both bosses are also fought in two phases.

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The mutated form of [[spoiler:Gregorio Silva]], and the source of the Red Mist separating Vein from the rest of the world. Juzo Mido forcibly introduces more Relics into the Successor's body so as to render him too insane to maintain the Gaol, and under the Queen's corruption [[spoiler: Gregorio]] is reduced to a bloodthirsty beast. [[spoiler: In all endings except for "Heirs", the lupine Lost summons the remaining unclaimed Relics throughout the land and absorbs them all in a last-ditch effort to preserve itself, transforming even further into a nearly-complete Queen.]]

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\n[[quoteright:350:https://static.tvtropes.org/pmwiki/pub/images/maxresdefault_1_005.jpg]]
[[caption-width-right:350:''The Skull King''/''Successor of the Brain'']]
[[quoteright:350:https://static.tvtropes.org/pmwiki/pub/images/maxresdefault_3_2_9.jpg]]
[[caption-width-right:350:''The Virgin Born'']]
The mutated form of [[spoiler:Gregorio Silva]], and the source of the Red Mist separating Vein from the rest of the world. Juzo Mido forcibly introduces more Relics into the Successor's body so as to render him too insane to maintain the Gaol, and under the Queen's corruption [[spoiler: Gregorio]] is reduced to a bloodthirsty beast. [[spoiler: In [[spoiler:In all endings except for "Heirs", the lupine Lost summons the remaining unclaimed Relics throughout the land and absorbs them all in a last-ditch effort to preserve itself, transforming even further into a nearly-complete Queen.]]



* BookEnds: The first Successor (Successor of the Ribcage) the Revenants fought is canine-like, the Skull King as the final boss is also canine-like.

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* BookEnds: The first Successor (Successor the party encountered was the Successor of the Ribcage) Ribcage, which looks like a wolf merged into the Revenants fought is canine-like, the body of a woman. The Skull King as the final boss is also canine-like.a werewolf-like creature.



* FinalBoss: Of the game.



* LightningBruiser: He moves at incredible speed and can easily destroy an unprepared player with his brutal attacks.
* NoSell: Both the Skull King and Virgin Born are immune to {{Standard Status Effect}}s and highly resist elemental damages. The Skull King is still vulnerable to Fire, Cold, and Lightning, but the Virgin Born resists all, even Blood.

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* FinalBoss: The Skull is the final enemy players need to defeat in order to complete the game. At least for the "Heirs" ending.
** TrueFinalBoss: As the Virgin Born. After the Skull King is defeated, [[spoiler:Silva]] loses control of the Relic of the Brain and draws in countless other Relics, resulting in an EldritchAbomination that [[spoiler:signifies the Queen is dangerously close to being revived]].
* GameplayAndStoryIntegration: A minor case, but the Skull King was originally a Revenant who utilized a Type Hound Blood Veil. As a result, he is the only Successor who uses their Blood Veil.
* LightningBruiser: He moves at incredible speed The Skull King is very fast, and can easily destroy an catch unprepared player players off guard if they aren't careful.
* MonsterProgenitor: Zigzagged. The Successor of the Blood was the first Successor to exist, but [[spoiler:Silva was the one who ordered the creation of other Successors,
with his brutal attacks.
him being the first and most important to undergo the process of becoming one]].
* NoSell: Both the Skull King and Virgin Born are immune to {{Standard Status Effect}}s and highly resist elemental damages. The While the Skull King is still vulnerable weak to Fire, Cold, Ice and Lightning, but Electricity, the Virgin Born resists all, even all elements, including Blood.



%%* OurWerewolvesAreDifferent:
* TrueFinalBoss: Kind of, and only applies to the Virgin Born. If going for an "Heirs" run, the game ends after the player has defeated the Skull King. In order to progress to and fight the Virgin Born, one must have fulfilled the conditions for at least the "To Eternity" ending.

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%%* OurWerewolvesAreDifferent:
* TrueFinalBoss: Kind of, and only applies to the Virgin Born. If going for an "Heirs" run, the game ends after the player has defeated OurWerewolvesAreDifferent: Ironic considering he's [[OurVampiresAreDifferent a vampire-like creature]], but the Skull King. In order to progress to and fight the Virgin Born, one must have fulfilled the conditions for at least the "To Eternity" ending.King looks like a hellish werewolf slash Cerberus monster whenever it uses its Blood Veil.
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* TheOneGuy: The only male of the successors that are fought, [[spoiler: until the player runs into the Skull King]].
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The mutated form of [[spoiler:Gregorio Silva]] that was turned into a Successor by Mido and slowly driven to insanity so that the Gaol of the Mist that [[spoiler:Silva]] maintained would be lifted and the Revenants could go outside and fight the Aragami. He mutates even further after absorbing even more Relics and turns into a giant monster.

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The mutated form of [[spoiler:Gregorio Silva]] that was turned into a Successor by Mido and slowly driven to insanity so that the Gaol of the Mist that [[spoiler:Silva]] maintained would be lifted Silva]], and the Revenants could go outside source of the Red Mist separating Vein from the rest of the world. Juzo Mido forcibly introduces more Relics into the Successor's body so as to render him too insane to maintain the Gaol, and fight under the Aragami. He mutates Queen's corruption [[spoiler: Gregorio]] is reduced to a bloodthirsty beast. [[spoiler: In all endings except for "Heirs", the lupine Lost summons the remaining unclaimed Relics throughout the land and absorbs them all in a last-ditch effort to preserve itself, transforming even further after absorbing even more Relics and turns into a giant monster.nearly-complete Queen.]]
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The mutated form of Gregorio Silva that was turned into a Successor by Mido and slowly driven to insanity so that the Gaol of the Mist that Silva maintained would be lifted and the Revenants could go outside and fight the Aragami. He mutates even further after absorbing even more Relics and turns into a giant monster.

to:

The mutated form of Gregorio Silva [[spoiler:Gregorio Silva]] that was turned into a Successor by Mido and slowly driven to insanity so that the Gaol of the Mist that Silva [[spoiler:Silva]] maintained would be lifted and the Revenants could go outside and fight the Aragami. He mutates even further after absorbing even more Relics and turns into a giant monster.
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* AppendageAssimilation: The Queen relics that they bear constantly corrode both their physical being and sanity, effectively turning them into new Queens once the assimilation is completed. Holding more than one relic inside one's body greatly accelerates this process.
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* SuperpoweredMook: They're Lost, but they're exceedingly powerful because [[SealedEvilInACan they've been grafted with pieces of Cruz's body.]]

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* SuperpoweredMook: They're Some of them are Lost, but they're exceedingly powerful because [[SealedEvilInACan they've been grafted with pieces of Cruz's body.]]body]].
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* {{Expy}}: The boss fights a lot like the Dancer of the Boreal Valley from ''Videogame/DarkSoulsIII'', walking on all fours, having fire moves accompanied by a flaming sword which gets used more in a second phase, and ambiguously sexy parts. [[spoiler: Similarly, both were the cause of cruel experimentation, the former by Juzo Mido and the latter by Pontiff Sulyvahn]]
* MsFanservice: Cat characteristics aside, she has a curvy physique, huge breasts and strikes sexy poses.

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* {{Expy}}: The boss fights a lot like the Dancer of the Boreal Valley from ''Videogame/DarkSoulsIII'', walking on all fours, having fire moves accompanied by a flaming sword which gets used more in a second phase, and ambiguously sexy parts. [[spoiler: Similarly, both were the cause results of cruel experimentation, the former by Juzo Mido and the latter by Pontiff Sulyvahn]]
* MsFanservice: Cat characteristics aside, she has a curvy physique, huge breasts and strikes sexy poses. [[spoiler: Emily Su's human form is also quite the eye candy herself, wearing a near skin-tight top with large breasts.]]
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* {{Expy}}: [[spoiler: the boss fights a lot like the Dancer of the Boreal Valley from ''Videogame/DarkSoulsIII'', walking on all fours, having fire moves, and ambigiously sexy parts.]]

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* {{Expy}}: [[spoiler: the The boss fights a lot like the Dancer of the Boreal Valley from ''Videogame/DarkSoulsIII'', walking on all fours, having fire moves, moves accompanied by a flaming sword which gets used more in a second phase, and ambigiously ambiguously sexy parts.]]parts. [[spoiler: Similarly, both were the cause of cruel experimentation, the former by Juzo Mido and the latter by Pontiff Sulyvahn]]
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* NoSell: Both the Skull King and Virgin Born are immune to {{Standard Status Effect}}s and highly resist elemental damages. The Skull King is still vulnerable to Fire, Cold, and Lightning, but the Virgin Born resists all, even Blood.


Added DiffLines:

* TrueFinalBoss: Kind of, and only applies to the Virgin Born. If going for an "Heirs" run, the game ends after the player has defeated the Skull King. In order to progress to and fight the Virgin Born, one must have fulfilled the conditions for at least the "To Eternity" ending.
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* NinjaPirateZombieRobot: A giant gas-masked CatGirl that also happens to shoot fireballs, be a super-agile predator, and also fights like a {{Ninja}}. Compared to the Successor of the Breath, this one doesn't exactly get an explanation [[spoiler: given that Emily Su was merely a Revenant soldier. At best, one could figure she was inspired by Yakumo's likes in an odd way.]]
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* {{Expy}}: [[spoiler: the boss fights a lot like the Dancer of the Boreal Valley from ''Videogame/DarkSoulsIII'', walking on all fours, having fire moves, and ambigiously sexy parts.]]
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[[folder:The Skull King/The Virgin Born]]

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[[folder:The [[folder:Successor of the Brain/The Skull King/The Virgin Born]]
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* {{Foreshadowing}}: As seen below in HumanityEnsues, she can be restored back to her original Revenant form. [[spoiler: Turns out that in the GoldenEnding, so can the other Successors once the Relics are finally pacified by the Protagonist once and for all.]]
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* XenomorphXerox: The Virgin Born's appearance has elements of a Xenomorph Queen.
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Removing spoilers in the descriptions since thats not allowed.


The Successor that awaits in the Cathedral of the Sacred Blood. A giant creature with a wolf-like head and four arms and two legs, it stands before the protagonist and company as if to block the door to the cathedral. [[spoiler: It's revealed that she is actually the scientist Aurora Valentino, a close friend of Karen, who volunteered to become a Successor to protect Karen who have also become a Successor.]]

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The Successor that awaits in the Cathedral of the Sacred Blood. A giant creature with a wolf-like head and four arms and two legs, it stands before the protagonist and company as if to block the door to the cathedral. [[spoiler: It's revealed that she is actually the scientist Aurora Valentino, a close friend of Karen, who volunteered to become a Successor to protect Karen who have also become a Successor.]]



The Successor who inherited Q.U.E.E.N.'s claws. It has deep red claws that shine like a crimson flame and two tails. Its head is fully masked, meaning that its facial expressions are unreadable, but the prey reflected in its eyes is undoubtedly invited to death. [[spoiler: It's revealed that she is actually Emily Su, who volunteered to become Mido's test subject, transformed into this Successor by one of the relics.]]

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The Successor who inherited Q.U.E.E.N.'s claws. It has deep red claws that shine like a crimson flame and two tails. Its head is fully masked, meaning that its facial expressions are unreadable, but the prey reflected in its eyes is undoubtedly invited to death. [[spoiler: It's revealed that she is actually Emily Su, who volunteered to become Mido's test subject, transformed into this Successor by one of the relics.]]



The Successor that awaits in the Ridge of Frozen Souls. A large animated suit of armor whose helmet is adorned with large antlers that awaits the protagonist and company among the frozen wastes. [[spoiler: It's revealed that he is actually Nicola Karstein, Mia's little brother who volunteered to become a Successor to protect his sister.]]

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The Successor that awaits in the Ridge of Frozen Souls. A large animated suit of armor whose helmet is adorned with large antlers that awaits the protagonist and company among the frozen wastes. [[spoiler: It's revealed that he is actually Nicola Karstein, Mia's little brother who volunteered to become a Successor to protect his sister.]]



The Successor that was recently sent into frenzy by Mido and awaits at the Crown of Sands. An angelic figure with a flower in one of its eye sockets and purple halo-like appendages sprouting out of her shoulder blade. [[spoiler:She holds Eva Roux's body in her chest.]]

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The Successor that was recently sent into frenzy by Mido and awaits at the Crown of Sands. An angelic figure with a flower in one of its eye sockets and purple halo-like appendages sprouting out of her shoulder blade. [[spoiler:She holds Eva Roux's body in her chest.]]



The mutated form of [[spoiler:Gregorio Silva]] that was turned into a Successor by Mido and slowly driven to insanity so that [[spoiler:the Gaol of the Mist that Silva maintained would be lifted and the Revenants could go outside and fight the Aragami]]. He mutates even further after absorbing even more Relics and turns into a giant monster.

to:

The mutated form of [[spoiler:Gregorio Silva]] Gregorio Silva that was turned into a Successor by Mido and slowly driven to insanity so that [[spoiler:the the Gaol of the Mist that Silva maintained would be lifted and the Revenants could go outside and fight the Aragami]].Aragami. He mutates even further after absorbing even more Relics and turns into a giant monster.
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* PosthumousCharacter: Not Eva herself, but the previous Successor of the Throat, who were killed by Jack just prior to the protagonist catching up with him in the Dried-up Trenches. His relic is then transferred into Eva, making her the new Successor.
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* WeaksauceWeakness: In defiance of all logic in a Soulslike, the best way to safely take on the Successor of the Ribcage is to get as close to her as possible. Specifically, she is mostly immobile throughout most of the fight, making it so that the player could sneak up behind the boss and [[GroinAttack literally smack her in the lady parts over and over again until she dies]]. She also has very little in the way of defense, as both of the attacks that could hit a player shanking her behind has obvious telegraphs that could simply be dodged out of the way in an instant.
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* PosthumousCharacter: Not Eva herself, but the previous Successor of the Throat, who were killed by Jack just prior to the protagonist catching up with him in the Dried-up Trenches. His relic is then transferred into Eva, making her the new Successor.
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* ReducedToDust: What every Successor ultimately becomes [[spoiler:if the protagonist cannot/does not save them, in addition to them dying. Despite all of them being rather large, the resulting ash pile is always tiny by comparison]].
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* BookEnds: The first Successor (Successor of the Ribcage) the Revenants fought is canine-like, the Skull King as the final boss is also canine-like.
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* ThemeMusicPowerup: Upon reaching the second phase of their respective boss fight, their original BattleThemeMusic goes away while "Heir of the Shingai" kicks in, signifying the Successor is now pulling out all the stops to kill you.

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* ThemeMusicPowerup: Upon reaching the second phase of their respective boss fight, their original BattleThemeMusic goes away while "Heir ''Heir of the Shingai" Shingai'' kicks in, signifying the Successor is now pulling out all the stops to kill you.
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* ThemeMusicPowerup: Upon reaching the second phase of their boss fights, their respective BattleThemeMusic goes away while "Heir of the Shingai" kicks in to signify the Successors are now pulling out all the stops to kill you.

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* ThemeMusicPowerup: Upon reaching the second phase of their boss fights, their respective boss fight, their original BattleThemeMusic goes away while "Heir of the Shingai" kicks in to signify in, signifying the Successors are Successor is now pulling out all the stops to kill you.
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ThemeMusicPowerup: Upon reaching the second phase of their boss fights, their respective BattleThemeMusic goes away while "Heir of the Shingai" kicks in to signify the Successors are now pulling out all the stops to kill you.

to:

* ThemeMusicPowerup: Upon reaching the second phase of their boss fights, their respective BattleThemeMusic goes away while "Heir of the Shingai" kicks in to signify the Successors are now pulling out all the stops to kill you.
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Added DiffLines:

ThemeMusicPowerup: Upon reaching the second phase of their boss fights, their respective BattleThemeMusic goes away while "Heir of the Shingai" kicks in to signify the Successors are now pulling out all the stops to kill you.
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* FinalBoss: Of the game.


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%%* OurWerewolvesAreDifferent:
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The mutated form of [[spoiler:Gregorio Silva]] that was turned into a Successor by Mido and slowly driven to insanity so that [[spoiler:the Gaol of the Mist that Silva maintained would be lifted and the Revenants could go outside and fight the Aragami]]. He mutates even further after absorbing even more Relics and turns into a giant monster.
----
* BloodyMurder: After he TurnsRed, his sword swings are followed by blood trails.
* DualWielding: With a {{BFS}} in each hand.
* EldritchAbomination: The Virgin Born form.
* LightningBruiser: He moves at incredible speed and can easily destroy an unprepared player with his brutal attacks.
* OneWingedAngel: After his initial defeat, [[spoiler:Silva]] pulls more Relics into his body and turns into the monstrous Virgin Born.
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* ShedArmorGainSpeed: He discards his shield and begins using his jetpack in the later portion of his fight.
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* NinjaPirateZombieRobot: An armoured knight with four eyes, a sawblade shield, moose antlers and ''a jetpack.'' Justified as it's the creation of [[spoiler: a little boy, who would find this sort of thing cool.]]
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* HumanityEnsues: Zigzagged. Unlike in the case of [[spoiler:Eva Roux]], who's transformation could be undone since she was only recently turned into a Successor, the rest are permanently stuck inside their new bodies. Their sanity can be restored if the protagonist purifies their Vestiges. [[spoiler:In the "Dwellers in the Dark" ending, the Successors' bodies are returned to normal]].

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* HumanityEnsues: Zigzagged. Unlike in the case of [[spoiler:Eva Roux]], who's transformation could be undone since she was only recently turned into a Successor, the rest are permanently stuck inside their new bodies. Their sanity can be restored if the protagonist purifies their Vestiges. [[spoiler:In the "Dwellers in the Dark" ending, the Successors' bodies are returned to normal]].normal, due to the fact that all the Relics were pulled out of their bodies, and thus ridding them of their transformed selves]].

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