- No one dinosaur species lived through the entire era, nor were they evenly distributed over the world.
- Hence, the dinosaurs didn't all live together in the same time and place.
- No humans lived contemporaneously with Mesozoic dinosaurs.
Dinosaurs and other extinct animals are divided here into four categories:
- Great Stock: *** What you think of when you think "Dinosaur"; have appeared everywhere in the media.
- Middle Stock: ** Have frequently appeared in media but not so often as the Great Stock. note
- Little Stock: * Have frequently appeared in documentary media but quite rarely in the more popular ones. note
- Non-Stock: Have appeared even more rarely in media (if at all). note See Useful Notes Prehistoric Life for these.
A brief history of popular depictions of dinosaursSee here for a more detailed article. 1850s: The Crystal Palace dinosaur sculptures in London introduced dinosaurs to the public. The image they provided was of scaly, bulky, doglike dragons (quite un-dinosaurian critters to our modern view). Introducing: Iguanodon and Megalosaurus. The park also introduced some non-dinosaur reptiles: the flying Pterodactylus and the swimming Mosasaurus, Ichthyosaurus, and Plesiosaurus. The 1864 novel Journey to the Center of the Earth portrayed the latter two as the first "antediluvian reptiles" ever in literature. Late 1800s: The excitement of the U.S. Bone Wars made dinosaurs interesting to the readership of newspapers and magazines that recounted the exploits and discoveries of Marsh and Cope. Stock dinosaurs found: Brontosaurus (Apatosaurus, though at the moment said synonymy is kind of rocky), Diplodocus, Allosaurus, Ceratosaurus, Stegosaurus, Triceratops, Trachodon (indeterminate hadrosaur), Hesperornis and Ichthyornis. Non-dinosaurs: Pteranodon, Elasmosaurus, Tylosaurus, and Dimetrodon. Early 1900s: Updated depictions of dinosaurs were brought to the general public by early paleoartists (beginning with Charles R. Knight), by distribution of skeleton casts which made life-sized and fairly life-like museum exhibits possible, and by dinosaurs being introduced to films. From this time on, dinosaurs and movie special effects were tightly coupled. Based on the finds during and since the Bone Wars, dinosaurs were now seen as a more varied bunch, with larger and... less large forms, bipedal or quadrupedal. They were still sluggish brutes destined for complete extinction, though. In 1940 Disney's Fantasia reached a large audience, but didn't change the media image much. Introducing: the aforementioned dinosaurs and non-dinosaurs from the Bone Wars plus Tyrannosaurus rex, Brachiosaurus (now split into Brachiosaurus altithorax and Giraffatitan brancai; the latter of which is infinitely more represented under the B. moniker), Styracosaurus, Ankylosaurus, Parasaurolophus, Corythosaurus, Plateosaurus, Ornithomimus/Struthiomimus, Protoceratops, and the non-dinosaur Rhamphorhynchus. 1970s: The Dinosaur Renaissance changed the image of dinosaurs to more active, more intelligent, more caring to their offspring, and well-adapted to their environment (and surviving extinction through bird descendants). Introducing: Deinonychus, and to some extent, Archaeopteryx (as a proper dinosaur; it's been well-known since the 19th century, but was considered a non-dinosaur before the Renaissance). 1980s/1990s/2000s: The Land Before Time and other works made at the end of the 1980s started popularizing the image of dinosaurs as set up by the Dinosaur Renaissance. Since the 1990s, scientifically up-to-date books and computer animation in films/shows (especially in the Jurassic Park and Walking with Dinosaurs franchises) have definitively completed the job. Introducing:
- By the Jurassic Park films and books: Velociraptor, Spinosaurus, Dilophosaurus, Gallimimus, Compsognathus, Carnotaurus, Maiasaura.
- By the Walking With docu series: Giganotosaurus, Argentinosaurus, Utahraptor, Therizinosaurus, and the non-dinosaur Liopleurodon.
- By the The Land Before Time films: Pachycephalosaurus and Microraptor.
- By others: Coelophysis, Oviraptor/Citipati, Baryonyx, Troodon/Stenonychosaurus, some alleged "biggest sauropods" (Supersaurus, "Ultrasaurus", "Seismosaurus"), and the non-dinosaur Quetzalcoatlus.
- Note: Some of the animals above were first discovered and described in the early 1900s (ex. Velociraptor, Spinosaurus) or even before (ex. Compsognathus), but they have ascended to true Stock only in the last few decades.
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Meat-eaters (usually)Most dinosaurs were herbivorous or omnivorous; the Theropod group contains all the carnivorous dinosaurs. Some of them were very small, while the biggest weighed as much as an elephant or a bit more, and were taller and much longer. Their legs were birdlike in structure; their feet had three main toes and usually a smaller reversed forth toe. All of them were bipedal; some had only tiny rudimentary forelimbs. Theropods are the only group of dinosaurs that has living members today, since they included the common ancestor of birds. Some close-to-bird theropods became omnivores and sometimes herbivores; the group includes the only toothless non-bird-dinosaurs. Many theropods are now known to have been feathered, but in films they are usually shown with lizard-like scales, while documentaries tend to be a bit wonky on the coverings of their theropods.
The King of the Cretaceous: Tyrannosaurus rex ***
Lived in western North America (between Alberta and Utah, and possibly as far south as Texas) 67-65 mya at the very end of the Age of Dinosaurs. Other tyrannosaurs lived in Asia in the same period, and other members of the tyrannosaurid family lived slightly earlier (still within the Late Cretaceous Period) in North America. note Together with Triceratops (and few others), Tyrannosaurus was one of the rare dinosaurs that was directly led to extinction by the asteroid/comet collision at the end of the Mesozoic Era. T. rex was discovered by Barnum Brown shortly before the start of the 20th century, and described by Henry Fairfield Osborn in 1905. Since then, it has been a hit with the audience and possibly the most famous dinosaur for almost a century. During this time depictions of T. rex have changed from the heavy, fat-bellied giant with goose-like gait and flexible tail seen in Fantasia to the slender, running beast seen in Jurassic Park. We were waiting to see it, or at least its chicks, depicted with feathersnote , and recently this idea has been seeping into pop-consciousness (thank you Dinosaur Revolution, Dinosaur Island, Pokémon, Mighty Magiswords, and Saurian). Despite only living for a couple of million years in a small part of the world, every visit to a dinosaur-populated time or place will have at least one T. rex appearing. This is for reasons better explained on the animal's own trope page. Yes, that's how big it is in media.
- Entry Time: 1905
- Trope Maker: Itself
Sickle-feet: Deinonychus, Velociraptor & Utahraptor, aka the "Raptors" ***
Raptors, or more formally dromaeosaurids, were bipedal carnivorous dinosaurs of the Cretaceous Period. They were small with long, thin tails and compact bodies. They were closely related to birds: their skeletal structure was bird-like, and since the late 1990s it has been proven they were also covered with pennaceous feathers. The most distinctive feature was the large, retractable "sickle claw" on their second toe. How it was used is being still debated. For decades, dromaeosaurids were depicted as hunting and attacking herbivores much bigger than itself, e.g. the classic Deinonychus hunting Tenontosaurus or even the five-ton Iguanodon. Like a pack of wolves, dromaeosaurids were envisioned attack their prey en masse, using their powerful claws to rend and to climb atop the herbivore. Since around 2000, dromaeosaurids have instead been suggested to have been mostly solitary hunters, taking prey the same size or larger than themselves, but leaving the very large ornithopods or sauropods alone (see Prehistoric Life). In the early 20th century, two small dinosaurs were discovered and described as generic small predators. Both were from the Late Cretaceous. While the finds were incomplete and difficult to interpret, we now know the animals were about 6.5 ft/2 m long and weighed about 33 lb./15 kg. Dromaeosaurus ("running lizard") lived in the Alberta region, while Velociraptor ("swift robber") lived in Mongolia and northern China 75-71 mya. For half a century, they were sorted away and largely ignored. Then... Deinonychus note ("terrible claw") was (re)discovered in 1964. It lived 115-108 mya in Early Cretaceous North America and was at the same time one of the largest and one of the earliest raptors, 11 ft/3.4 m long and weighing 160 lb/73 kg. Even though some illustrations showed it as tall as an adult man, it would actually only reach his hips if alive today. Some years later, more complete remains of Velociraptor were found, showing that it was similar to Deinonychus but even smaller (the weight of a large turkey or an ocelot). Deinonychus was described by John Ostrom in 1969 in an influential monograph that kicked off the Dinosaur Renaissance. After that, paleontologists, especially Ostrom's pupil Bob Bakker, began to debate if the traits ascribed to Deinonychus (agility, smartness, warm-bloodedness, social behavior) should be extended to all dromaeosaurids, or possibly to all theropods, or even to all dinosaurs. In the 1980s, one paleontologist (Gregory Paul) claimed that Deinonychus and Velociraptor were actually the same genus and that the species Deinonychus antirrhopus should be renamed "Velociraptor antirrhopus"; author Michael Crichton picked up this idea, showing both Deinonychus and Velociraptor proper in his Jurassic Park novel as distinct species within one single genus, "Velociraptor". note Works from before the 1970s never represent dromaeosaurids, simply because they were scientifically too obscure at the time. Significantly, between 1970 and the Jurassic Park mania in the 1990s, the most represented "raptor" (though not yet known by that term) in popular culture was the biggest known at the time, Deinonychus, while the less-impressive Velociraptor was totally unknown to laymen (not counting the antecedent dino-fans). For instance, see Dino-Riders, Carnosaur, the RuneQuest Borderlands tabletop RPG adventure, or even the Dutch metal-band named Deinonychus. It was Jurassic Park that apparently caused Velociraptor to displace Deinonychus as the stock sickle-clawed dino (even documentary media started showing the "veloci" more often thanks to the film), and started the usage of "raptor" for dromaeosaurid in the mind of the public. (Prior to this, "raptor" was used only to indicate birds of prey.) Without Jurassic Park, Velociraptor would have almost-surely remained in the Non-Stock realm forever. There are several issues with the depiction of raptors in the film. At the same time that the name Velociraptor became popular, a new dromaeosaurid was discovered in Utah. This animal was even larger and slightly older than Deinonychus, living 128-105 mya and being 23 ft/7 m long and as tall as a human. It was named Utahraptor, beginning an awesome case of science culture Ascended Fanon — before Jurassic Park, no genus of dromaeosaurids except Velociraptor had the -raptor suffix to its name. Since the film, paleontologists started to use it for naming most new dromaeosaurids. Despite the scantiness of its remains, the discovery of Utahraptor was much reported in media, as it incidentally matched the size of the oversized JP raptors (or rather, was even longer than they were). Many then reported the Utahraptor as "the most fearsome killing-machines of all times", capable of killing, in packs, the biggest sauropods and even of destroying entire dinosaur species. However, Walking with Dinosaurs was not so extreme, showing Utahraptor hunting the relatively smaller Iguanodon — not in Utah but in Europe for some reason. Utahraptor, however, has proven to be quite the oddball for a dromaeosaur since it's rise to stardom, with new remains revealing it to have had stumpy arms, a short tail, short, robust legs and a unique dentition, similar to animals like Masiakasaurus. What exactly this bizarre new look entails for the behavior and ecology of Utahraptor, as of now, remains uncertain. Speaking of misplaced wildlife, you can expect any of these three to be placed in the same habitat as at least Tyrannosaurus and Triceratops. In reality, Deinonychus and Utahraptor were already extinct by the time T. rex came along and Velociraptor lived on the other side of the planetnote . However, this inaccuracy was vindicated somewhat by the discovery of Acheroraptor (described in 2013) and Dakotaraptor (described in 2015), which greatly resembled Velociraptor and Deinonychus respectively, and the latter was roughly the size of Utahraptor. Contrary to popular depictions, actual raptor hunting tactics did not involve high-speed pursuits or using the claws to disembowel prey. What they did involve, however, was a ridiculous amount of Nightmare Fuel. Dromaeosaurs could only run at about 45 miles/hour for the faster species (Deinonychus and Velociraptor) and 20 miles per hour for the larger, more robust Utahraptor (this is still faster than any human), but they had Super Reflexes and superb Combat Parkour skills, able to leap, flip, dodge, and accelerate with shocking ease. They were good climbers, and their feathered wings probably helped them control their falls as well as muffling sound. The talons provided a near-unbreakable grip, and were used as grappling devices and stabbing weapons. Recent research into their eye sockets prove they had superb vision and were able to see very well in the dead of night, which was when they would use Jump Scare tactics to sneak up on unwary prey. Put together, they provide a frightening picture of an agile, stealthy, powerful hunter stalking victims at night from the treetops, completely silent and invisible, before leaping in as Death from Above, dodging counterattacks using Combat Parkour, latching on with a formidable grip, and either stabbing the prey in the neck or pinning it down and eating it alive as it writhed in agony.
Post-rex stock theropods
Since the '80s/'90s, a handful of large theropods have started to filter into pop-consciousness, often after a single remarkable appearance in Movieland: Spinosaurus (Jurassic Park III), Giganotosaurus, Carnotaurus (Disney's Disney/Dinosaur), and Baryonyx (Ice Age 3: Dawn of the Dinosaurs). Despite having some cool traits (crests, horns, claws, or sheer size), none of them has managed to replace T. rex as the "King Dinosaur" — at least for now. Though relatively small, Dilophosaurus (popularized by the 1st Jurassic Park film) is here for comparison.
Spiny-backed River Monster: Spinosaurus **
Lived in Northern Africa 112-97 mya, during the Cretaceous Period. At present, this is the biggest theropod; no other matches it in bulk and length. Spinosaurus is one of the most recognizable theropods with its 5 ft/1.5 m-tall spines on its back. In the most common interpretation the spines form a "sail" similar to that of the non-dinosaur Dimetrodon. Some suggest that they instead supported a ridge, while others thought they came from another dinosaur altogether. A sail could have been useful as a thermoregulating device to prevent overheating and/or as a display tool, and a ridge could have been for display, making the animal seem larger, as well as storing extra energy gained from the giant fish and other prey that Spinosaurus fed on. Spinosaurus was first described in 1915 by a German paleontologist, but its remains are very scanty: its skull is incomplete, and we have no limb bones. The best spinosaur find was stored in a German museum, which was destroyed by aerial bombing during World War II. In older drawings Spinosaurus had a head like a generic "carnosaur"; today it is generally accepted that its head was similar to a crocodile's. Due to the fragmentary nature of its remains, the actual overall size is in debate; it was once thought the same length of an average Tyrannosaurus (40 ft/12 m), but many paleontologists wanted to set the length at 50 ft/15 m. Lack of real evidence for this left T. rex with the official record until the discovery of Giganotosaurus in the mid-1990s. Meanwhile, the spinosaur remained an only-known-among-dino-lovers dinosaur. Then, in the year 2001... ... Jurassic Park III (which fans don’t really like to talk about but was nonetheless popular) changed this situation in a blink. This film introduced the spinosaur to the audience as "bigger and badder" than a Tyrannosaurus rex, and easily capable of defeating the latter in a fight. Many dino-fans complained that the JP spinosaur was oversized and altered to make it a sort of Pseudo-Rex thing. Then, new discoveries told us Spielberg wasn't totally wrong: Spinosaurus really was bigger than Tyrannosaurus rex, in fact larger than shown in the film. Not only that, it was indeed the biggest of the lot, and is still considered to be. There is some controversy regarding Spinosaurus’ diet and way-of-life: did it prey on fish like its smaller cousin Baryonyx (see below), or on giant herbivores like Tyrannosaurus did? Experts tended to prefer the first option at the time Jurassic Park III was produced, and this fostered even more criticism about the film portrayal as the Ultimate Superpredator. Today Spinosaurus’s lifestyle is generally believed to have been midway between these two extremes: an opportunist like a giant, clawed, saltwater crocodile, attacking other smaller dinosaurs when given the opportunity, as well as eating giant fish (mostly sharks and other fish the size of most dinosaurs) and possibly crocodiles, a feat requiring tremendous levels of strength, and using its size to steal kills from other predators. We're unsure about the latter, though: Carcharodontosaurus was specially adapted to big-game hunting and could open its jaws very wide to inflict severe slicing cuts, which could likely cause the spinosaur to bleed to death if the two fought. Spinosaurus’ large size would, however, make it a hard target to bite for other predators. In 2014, Spinosaurus received an almost-total makeover after new fossils were discovered. In addition to tiny pores in its skull that might have enabled it to sense underwater prey (which was already known by then), Spinosaurus also had a relatively small pelvis and short hind legs with flat — possibly webbed — hind feet, among other adaptations for a semiaquatic lifestyle — all in all showing a surprising degree of convergent evolution with the ancestors of modern whales. Unrelated to its amphibious makeover, its reconstruction also gives it a dip in the middle of its dorsal spines similar to that of its relative Ichthyovenator. It has been suggested that the hind limbs were scaled incorrectly but then the people who made the discoveries have responded putting these supposed corrections to doubt. Also, the tall spine on the sail might have actually been located further back on the body thus forming a more shallow and gradual version of the "classic" Spinosaurus sail note , although this too is debatable. Finally, it has been suggested that Spinosaurus held its neck vertically rather than horizontally. This would have shifted the center of gravity back and allow Spinosaurus to walk bipedally albeit in a tripodal stance.note Other alternatives for land locomation include hobbling like a pangolin using its forelimbs to prop itself back up, or maybe even crawling on its belly like a crocodile or a pinniped. Time will tell if the legs were really that short, although it does seem to have had shorter legs (but seemingly still a biped) and been almost entirely aquatic.
Giant of the south: Giganotosaurus *
Giganotosaurus (NOT "Gigantosaurus"; that name was used for an invalid sauropod) lived in Late Cretaceous South America 97 million years ago. A close relative of Allosaurus, it had a bigger head (6 ft/1.80 m long, even longer than a Tyrannosaur’s) and a stockier build: its looks seems rather like a cross between an allosaur and a tyrannosaur — incidentally, making the classic hybrid allo/tyranno so often seen in classic films (Fantasia, 1 Million B.C.… ) a sort of Truth in Television. Discovered in 1993 and officially described two years later, Giganotosaurus was celebrated as "the biggest predatory dinosaur ever," surpassing Tyrannosaurus, of whom the largest specimen known (the famous Sue) was discovered a few years before. The "giga" remained the record-holder until new Spinosaurus fossils were discovered in the 2000s, and the re-examination of the descriptions of older finds, reminded us that the latter was even larger, something already postulated but ignored for 80 years. At the same time, re-examination of Giganotosaurus remains show an animal not much larger that Tyrannosaurus; the only advantage in length is due to a longer snout, and were the two animals placed side-by-side, they'd appear to be the same size. Its close relative Carcharodontosaurus (known since the first half of the 20th century in the form of teeth, but rediscovered in 1996), got the same treatment in the 1990s, ultimately lost the struggle for widespread recognition; and both ended up overshadowed in popular culture by Spinosaurus. Giganotosaurus remains one of the most powerful meat-eaters that ever lived; and it's just starting to gain popularity. The fact that it could have possibly hunted some of the largest sauropods — possibly including of the utterly vast Argentinosaurus — means that it may become very popular in the future. If that doesn't sound cool enough, then consider that to do so, it would have had to be a pack hunter. Walking with Dinosaurs did a special on just how badass such a hunt would be (even though in the show the Argentinosaurus that became prey was a juvenile). Though there isn't any evidence for pack behavior in Giganotosaurus, there might be for its relative, the recently-discovered Mapusaurus, which was the same length but had a more slender frame.
Meat-loving, sprinting bull: Carnotaurus *
Another South American theropod like the former, Carnotaurusnote lived in the Late Cretaceous in a younger age, 70 million years ago. Discovered only in 1985, it is known from a single specimen, but this was one of these things every paleontologist wishes to find: one of very few big theropods so far found with skin impressions. As these prints are from the whole right side of its body, Carnotaurus is one of the only large dinosaurs whose external look is known with a reasonable degree of certainty, together with the so-called "hadrosaur mummies" (see "Hadrosaurs"). We don't know what was the coloration of the living animal, however. But this is not all: our carnotaur has revealed to be one of the strangest-looking dinosaurs known. Forelimbs even tinier than those of T. rex, sort of useless stubs with no true fingers (though three clawed fingers appear in many portrayals, and sometimes even a small Iguanodon-like thumbclaw for each hand) that contrast vividly with the long legs apt for high-speed runs note . Unusually shortened head (some compare it to a bulldog's). Above all, a couple of unique bull-like horns above the eyes which no other known theropod had (Carnotaurus means "meat-[eating] bull"). Finally, its skin was covered by rows of horny tubercles. The horns and the skin make Carnotaurus a quite dragon-looking dinosaur. Ironically, with its slender body, tiny forearms, and fragile lower jaws, it's hard to imagine how it could kill large prey in Real Life, esecially if you think it doesn't come close to rivalling Tyrannosaurus in size (it was only about 22 ft/7 m or so in length, while a big rex would be about 43 ft/13 m). A clue may come from the fact it is the fastest non-bird dinosaur ever discovered, being able to run at 75 miles per hour. It has been suggested it ran into prey at full speed with jaws open, using its head as a sledgehammer. However, Carnotaurus has become somewhat popular in the last decades, thanks to its striking look. It was portrayed as an oversized, pseudo-rex villain in the 2000 Disney’s Dinosaur. Here, the biggest carnotaur appears even larger than a spinosaur, able to lift an Iguanodon with its jaws, and fling it to death against a rock. In Real Life the 1-ton Carnotaurus was much smaller and weaker than the 5-ton Iguanodon, and lived several million years after the latter. Before that, it also showed up in Michael Crichton's second Jurassic Park book, where its size was portrayed more accurately, but to up the threat level, it was given (quite implausible) chameleon-style stealth abilities. Note that neither modern birds nor crocodilians can change their colors like chameleons do. Carnotaurus may be the responsible for the recent decline of the classic carnivore Ceratosaurus in media, as both dinosaurs had a similarly horned/tubercled look and the two dinos might be confused with each other, even though their look was rather different (see also Ceratosaurus in another section). But also note a bit of resemblance both in shape and in name between the carnotaur and a mythical critter, the Minotaur. This association may have at least subconsciously led to it becoming a go-to bad guy dinosaur.
Heavy claws: Baryonyx *
A cousin of Spinosaurus that lived in Europe in the Early Cretaceous, 130-125 mya, alongside Iguanodon. Discovered in 1983 in Southern England and named in 1986, its find got massive media coverage at the time (especially in the British media); in part because, being 30 ft/9 m long, Baryonyx was the largest and most complete European giant theropod, but mostly because it was very different from other dinosaurs known at the time, with its crocodile-like jaws lined with an incredible 96 teeth note (Spinosaurus was still portrayed with a token-theropod head in the 1980s and '90s), and very special forelimbs. Baryonyx means "heavy claw" and the animal has been nicknamed "Claws" because of its 10-inch/25-cm hook-like thumb-claws, bigger than the other two fingers on each hand. We don't know if Spinosaurus had these hook hands as well. Baryonyx’s forelimbs were longer and stronger than in most other theropods, but the structure of the forefeet seems to preclude quadrupedal walking (contrary to what is sometimes shown in illustrations); it is speculated, however, that Baryonyx might have fed by resting on its front legs on a riverbank and sweeping large fish such as the carp-like Lepidotes from the river with its powerful claw, a bit like grizzly bears do with salmon. We know for sure fish were included in its diet: scales of Lepidotes were found inside the ribcage of the only well-known Baryonyx specimen. Baryonyx was the first discovered fish-eater among dinosaurs, and several traits scientists today assign to Spinosaurus were initially based on Baryonyx. Together, these dinosaurs (plus Suchomimus and few others) form the spinosaurid family. However, Baryonyx was quite different from Spinosaurus: it had no sail on its backnote , and was considerably smaller (10 m long and weighing 2 tons, like an Allosaurus). Its head was thinner with a small bump on its top, and gharial-like jaws with twice the teeth of most other theropods. Baryonyx was less aquatic than Spinosaurus: fish might have made a greater part of its diet, possibly with occasional carrion and small land animals as a supplement. Its short hindlegs show it was not an especially fast runner; moreover, its blunt croc-like teeth and weak thin jaws probably prevented the "bary" to kill prey the size of a fully-grown Iguanodon in spite of the former's huge thumbclaws (incidentally, Iguanodon too had oversized thumbnails, but they were almost-straight and not curved like the carnivore's). Since the 1980s, "Claws" has been one of the most frequently-portrayed large theropods in popular dino-books. On the other hand, it has long been ignored in Fictionland and even most TV documentaries. It came into the spotlight only in 2009 thanks to the Ice Age 3: Dawn of the Dinosaurs film. Here an oversized Baryonyx called Rudy is the pseudo-rex Big Bad who is even bigger than the ''JPIII'' spinosaur or the real-life one. The baryonyx is unnamed however (some dino-fans wrongly think he's a Suchomimus), and also quite inaccurate, with a head shaped like a literal crocodile's and hands lacking the distinctive thumbclaws. Rudy is an albino; unlike chameleon-like carnotaurs, albino dinosaurs were possible in Real Life but probably very rare.
Double crest: Dilophosaurus **Dilophosaurus lived 197 million years ago in Early Jurassic North America. It was one of the first theropods to have exceeded human size (20 ft / 6 m long and weighing 500 kg). Smaller and more slender than the other carnivores listed above, it was a fairly close relative of the dog-sized Coelophysis. Its most easily recognizable trait is the two parallel crests on its skull (perhaps occurring only in males), probably used for display. These fragile and vulnerable structures indicate that it was no badass dinosaur. Its head was long and narrow with weak jaws and teeth, and the upper jaw also had a deep indentation on each side near the tip of the snout, making the whole structure even weaker.note Scientists have usually said the dilophosaur was a mere scavenger or a small prey hunter, even though some had initially described it as the "the first giant killer dinosaur". It was indeed one of the biggest terrestrian carnivores of its time (some docu-portrayals show it as a rather unlikely powerful super-predator); however, really powerful meat-eating dinosaurs only started to appear in the Middle Jurassic, for example Megalosaurus. note Dilophosaurus was first described in 1954 in Arizona from scant remains lacking the head, and was initially thought to be another species of the "wastebasket taxon" Megalosaurus. The first head complete with double crest was found only several years later; in 1970, the animal received the name Dilophosaurus, "double-crested lizard". Several footprints found in Early Jurassic terrains of the USA might have been made by dilophosaurs, but the exact identity of dinosaurian tracks cannot be told with certainty. One of these tracks was made by a swimming animal which barely touched the bottom with the tips of its feet. It's unlikely that many people outside the dino-fandom had ever heard of Dilophosaurus before the novel Jurassic Park was published in 1990. Here it was depicted as capable of spitting venom like some species of cobra, which it probably couldn't do in Real Life: venomous saliva is unknown among modern birds and crocodiles. Two years later, the JP movie made the dilophosaur even more popular and even more incorrect. Its size was greatly decreased note , but above all, Spielberg added a totally improbable Frilled Lizard-like cowl on its neck. It certainly did not have this frill; it would require a lot of specific musculature on the neck, and the imprint of this would be visible on the skeleton (it isn't). Still, most later popular depictions have represented Dilophosaurus with this thing. Just like Velociraptor, Dilophosaurus became a household name after the film, commonly known as the Spitting Dinosaur. Even though it has not appeared in any of the sequels (except for a quick cameo as a hologram in Jurassic World), the JP portrayal has remained in pop-consciousness, coincidentally preventing the Real Life animal from become more widely-known. Today, the ever-increasing public interest in dinosaurs (mainly started thanks to Jurassic Park) is making Spielberg’s Mix-and-Match Critter more and more of a Lost Subtrope.
Pre-rex stock theropods
Some large meat-eaters entered pop culture before T. rex: Allosaurus, Ceratosaurus, and Megalosaurus. But they have been less frequently portrayed since T. rex was discovered.
The King of the Jurassic: Allosaurus **
Allosaurus lived 155 to 150 million years ago in North America, with some fossils found in Europe and maybe Africa. Along with Tyrannosaurus, it has traditionally been the large carnivorous dinosaur. Allosaurus is the scientifically most well-known large theropod: dozens of specimens have been found so far in Western USA, including a veritable "graveyard" in Utah.note Many young individuals are also known. First discovered in 1877 during the Bone Wars, Allosaurus literally means "other lizard" or "strange lizard", but Othniel Charles Marsh's article naming it gives no reason for the bland choice. The most well-known species is Allosaurus fragilis ("the other fragile lizard"); some fragmentary remains of unusually large size are often classified in separate genera (see Prehistoric Life). Some scanty fossils from Early Cretaceous Australia used to be classified as a small-sized late-surviving Allosaurus species, but were reclassified in 2009 as a totally different theropod, Australovenator. Even scantier remains were found in the USA before Allosaurus was officially described in 1877; they were labeled Antrodemus, but possibly belong to Allosaurus as well. If true, the former might become the valid name for this dinosaur. note Allosaurus was the top predator in the Late Jurassic, sometimes referred as "the tyrannosaur of the Jurassic". Its hunting behavior is still uncertain: we're not sure if it was mainly a pack-hunter or a solitary ambush-predator. In documentaries and pop-books it usually appears as a pack-hunter capable of bringing down the biggest sauropods like Diplodocus (like in the memorable The Ballad Of Big Al), Apatosaurus, or even Brachiosaurus. Alternatively, it is shown in a battle against the armored Stegosaurus (the Jurassic equivalent of the Tyrannosaurus-vs-Triceratops Cretaceous duel). All this might be Truth in Television since all these animals lived together in North America in the same period, but more probably Allosaurus more often hunted easier prey such as young sauropods, young stegosaurs, and ornithopods like Camptosaurus. There are, however, stegosaur fossils showing Allosaurus bite marks and Allosaurus fossils that show wounds created by stegosaur tails. Allosaurus entered pop culture before Tyrannosaurus. After its description, it was briefly considered the "biggest land carnivore ever" together with Megalosaurus. In Conan Doyle’s The Lost World (1912) the two scientists encounter a giant carnivore, and argue about whether it is an Allosaurus or a Megalosaurus (maybe a reference to the then-recent Bone Wars). Soon afterward, both dinosaurs got overshadowed by the more impressive (and much cooler-named) Tyrannosaurus rex in pop-media, especially cinema. Allosaurus has somehow managed to survive the supremacy of the rex... automatically becoming its Poor Man's Substitute, as the two animals tend to be easily confused with each other in the public mind. Actually, Allosaurus is rather easy to distinguish from T. rex. It was generally smaller (the classic species was slightly shorter and about one half of the weight of a T. rex), had shorter legs, a longer tail, slimmer body and neck, narrower head, weaker lower jaw, smaller teeth, and a pair of "bosses" in front of its eyes (maybe covered in keratin in Real Life, making them like small "horns"). Above all, it had longer front arms with three clawed fingers rather than two. Sadly, all these differences tend to be glossed over in popular media. The fact that T. rex itself has often been depicted with long arms with three functional digits (e.g. in Disney's Fantasia) doesn't help, either. Among the official Allosaurus appearances in cinema, the Ray Harryhausen ones are the most remembered. The allosaur is the go-to Big Bad of his movies, appearing in One Million Years BC and playing the role of Gwangi in The Valley of Gwangi. Ray's critters looked just like that of Fantasia, with the same mishmash of allosaur and tyrannosaur features (and with the same outdated erect body, serpentine tail and goose-gait); the only difference is that Harryhausen's theropods have Evil Eyebrows (this may be forgivable for some, considering the aforementioned eye bosses).
Horned rex, or just underdog?: Ceratosaurus *Ceratosaurus lived in the same place as Allosaurus in the Late Jurassic, 153-148 mya. Usually 17-23 ft/5-7 m long, it was smaller than most other Stock Theropods above, but still a powerful animal. Its look was like that of an undersized allosaur, with the same eye-bosses and long forelimbs. Its name, "horned lizard", underlines its more distinctive anatomical feature: a laterally flat crest on its nose, classically described as a "nasal horn". note It was also the only known theropod to have armor in the form of bony plates along the middle of its back. While the "horn" is a constant in Ceratosaurus portrayals (don't be surprised to see it shaped like a rhino's), the armor can be left out altogether, or alternatively, modified to make the animal similar to a dragon. Despite its appearance, Ceratosaurus was actually more archaic than Allosaurus. Allosaurus belongs to the tetanuran branch of theropods, while Ceratosaurus is the namesake of its own branch, ceratosaurs. The latter can be told apart from tetanurans by the primitive shape of their pelvis, more flexible tails, and a remnant forth finger on each hand (tetanuran theropods never have more than three fingers). Most of the other theropods discussed here are tetanurans, except fellow ceratosaur Carnotaurus, and the more primitive Dilophosaurus and Coelophysis. Ceratosaurus was first found during the Bone Wars like Allosaurus, but is much rarer in the fossil record than the latter: many paleontologists suspect it was more solitary than allosaurs. In paleo-art and documentaries, Ceratosaurus is usually shown either as a scavenger or an underdog predator, but may occasionally be shown as a pack-hunter of big game. While Allosaurus is seen as the "lion" of its time, Ceratosaurus might be considered the "hyena"; with its smaller size, longer teeth and stronger jaws, the comparison works. Since Real Life spotted hyenas are not lions' underdogs (as seen in The Lion King) with both co-dominating the top-predator niche, its possible that ceratosaurs and allosaurs had a similar relationship. The horn on its nose and the armor make Ceratosaurus the most "dragon-looking" of the theropods known at the start of the 20th century. It's not a big surprise that it appeared in so many classic dino-films, from simple cameos (like Fantasia) up to being the main dino-actor (like the Ray Harryhausen film Animal World, in which two ceratosaurs get into a fight and fall off a cliff). Ceratosaurus holds the record of being the first dinosaur ever shown in non-animated cinema — the 1914 film Brute Force pitted cavemen vs dinosaurs and started the Dinosaurs Are Dragons trope. In later fiction Ceratosaurus received the same treatment as Allosaurus, acting as a T. rex substitute for the Big Bad part. With its distinctive look, Ceratosaurus is less likely than Allosaurus to be confused with Tyrannosaurus; on the other hand, its size is often exaggerated to make it more of a "horned tyrannosaur". Ceratosaurus is quite rare in films these days: the only recent example is a short cameo in Jurassic Park III, in which it's not even named (but at least is correctly sized). Even modern documentaries rarely represent it — the Walking with Dinosaurs series didn't show it at all. The recent Ceratosaurus decline is probably due to the occurrence of other, newly-discovered theropods since the '70s: Carnotaurus in particular, being similar yet even more badass looking.
- Entry Time: 1914
- Trope Maker: Brute Force (film)
The first named non-bird dinosaur: Megalosaurus *
We've already mentioned Megalosaurus more than once. Why? Well, both because it was the first giant theropod known to science, and because shows neatly how Science Marches On is normal stuff in dino-science. Its first remain, the extremity of a leg-bone found in 1676 in England near Oxford, was mistaken by Robert Plot for the remain of an ancient giant man (others named this remain Scrotum humanum because of its shape), but this fossil has since been lost. Later, a half lower jaw with a single tooth left was found in 1824 in Southern England; its discoverer, reverend and geologist William Buckland, described it as belonging to a "big lizard" (the meaning of its name, which started the tradition of "saurus" in dinosaur names). Buckland didn’t realize he had named the very first non-avian dinosaur. At the time, the Dinosaur category didn't even exist in scientific literature. The scientific and popular view of what a megalosaur was has gone through several drastic changes. The first attempt at reconstruction, the life-size sculpture in Crystal Palace Park constructed in the 1850s, made the Megalosaurus a dragon-like animal walking on all fours. Next to the Megalosaurus was an Iguanodon sculpture (also quadrupedal), and for several decades this was the stock image of the world of dinosaurs: one herbivorous dragon facing a carnivorous dragon in combat◊. From this time is Charles Dickens' novel Bleak House (1853) where Megalosaurus is mentioned, described as an "elephantine lizard". New genera of large carnivores were described during the Bone Wars, such as Allosaurus and Ceratosaurus, whose more complete remains showed clearly bipedal animals. Since then, Megalosaurus has also been reconstructed as bipedal. Even with their correct shapes, the "Megalosaurus vs Iguanodon" battle has remained a classic in non-fictional portrayals (a bit like "Tyrannosaurus vs Triceratops"), even though in Real Life the megalosaur was a middle Jurassic animal (166 mya), while the iguanodont lived 40 million years later in the Early Cretaceous. The tendence to classify theropod fossils of every kind as Megalosaurus started soon after its first description. After the Bone Wars, Megalosaurus still remained a "Wastebasket taxon" to which all finds that were too incomplete or too ambiguous were assigned. Megalosauruses cropped up everywhere from North America to Africa and from Early Jurassic to Late Cretaceous. Finally, scientists sorted out outsiders into more than 20 genera (Carcharodontosaurus, Dilophosaurus, Eustreptospondylus, Majungasaurus, and Proceratosaurus among them). This cleanup has yet to be definitively finished. The only-valid Megalosaurus is a fairly generic theropod some 30 ft/9 m in length, similar to an elongated allosaur but smaller and more primitive. Even though its historical relevance makes it a common sight in classic and modern dino-books, the "big lizard" didn't go a long way in popular works after the two important mentions in early literature (Bleak House and The Lost World). In the 20th century it heavily suffered the competition with Tyrannosaurus and Allosaurus — and the resolution of the "wastebasket" issue didn't make its case better. Apart from some occasional documentary, you have little chance of seeing any megalosaur either in cinema or in TV media — just as an example, Walking with Dinosaurs chose to portray the contemporary close-relative Eustreptospondylus in the Jurassic Europe episode. There is, however, the curious case of the TV show Dinosaurs, which has one "megalosaur" in the form of Earl Sinclair: but he doesn't look particularly like any dinosaur at all.
- Entry Time: 1852
- Trope Maker: Crystal Palace Park
Smaller Tyrants: Albertosaurus *Albertosaurus is the most abundant tyrannosaur in fossil record, and also the second big-sized theropod by wealth of fossil material, just after the unbeatable Allosaurus. And yet, Albertosaurus has not gained much attention in films and comics as Tyrannosaurus — tyrannosaurids are so similar to each other that if one appears in cinema, people will always call it T. rex. To compensate, Albertosaurus is a very common sight in many paleo-books, just as common as several great-stock theropods. Naturally, it is portrayed as the superpredator of its time, North America 80-75 million years ago, 10 million years before T. rex. The menu of an Albertosaurus was probably not monotonous; several kinds of herbivores roamed North American plains at the time, from ceratopsians to hadrosaurs, from the armored ankylosaurs to small swift hypsilophodonts and ornithomimids. Even though tyrannosaurids are classically shown battling some powerful prey, they more probably hunted young individuals more often, to avoid the risk of fatal injuries or consequent infections. Compared with the legendary Tyrannosaurus rex, Albertosaurus was like a leopard compared with a lion; smaller (25 ft/7.6 m long against the 40 ft of T. rex), it was also more slender, with longer, thinner jaws, smaller teeth, and more agile legs apt to higher top speeds than Tyrannosaurus. Even the herbivores which shared their world were matched with these predators; those which lived alongside T. rex were bigger, slower and more heavily armored than those living with Albertosaurus. Albertosaurus was also the first dinosaur ever discovered in Canada, at the end of the 19th century, but was named only in 1905 (incidentally, the same year as Tyrannosaurus) after the Canadian province of Alberta, where most of the abundant Canadian dinos have been discovered. Albertosaurus has also contributed indirectly to the popular image of tyrannosaurs. The forelimbs of Albertosaurus have been known since its very first find, while those of T. rex were first discovered only in the 1990s; for almost a century the well-known two-fingered hands of "rex" have been modeled upon those of Albertosaurus, debunking at the time the old pop-cultural Hand Wave about portraying three-fingered tyrannosaurs. note
Other large theropodsSorry, these ones aren't here. If you're looking for Tarbosaurus, Megaraptor, Suchomimus, Majungasaurus, and others, see here.
Bird-like / small theropods
Several bird-like and/or small theropods make appearances in media, although less commonly than the iconic "raptors".
Dino-like bird or bird-like dino?: Archaeopteryx, aka the "Missing Link" **
Archaeopteryx note lived around 150-148 mya in Late Jurassic Europe. Its name means "ancient wing" or "ancient feather"; another obsolete synonym very common in old textbooks was Archaeornis, "ancient bird". Both terms are very meaningful about its historical relevance. It is sometimes known as the "Urvogel", which is German for "original bird". Archaeopteryx was discovered 1861 in the famed Solnhofen deposit in Germany, whose rocks have preserved fossils so well that even soft parts of animals are visible. Because of this, most specimens of Archaeopteryx found later in Germany were found with impressions of feathers. Charles Darwin had published On the Origin of Species two years earlier, and in the following debate this "half-reptile, half-bird" became a key piece of evidence, as the perfect example of a "missing link" between two animal classes. A century later, Archaeopteryx was again used as crucial evidence, this time in John Ostrom's theory that modern birds had evolved within the theropod group. Before that, most scientists considered Archaeopteryx and its bird-descendants only distant relatives of dinosaurs, in part because in traditional zoology Feathers = Bird, and no other dinosaur was known with imprints of feathers. Even though one scientist (Darwin's pupil T.H. Huxley) already recognized the dinosaurian origins of birds as early as the late 19th century (by studying the skeletal features), this was largely accepted only after the Dinosaur Renaissance note and definitively proven only in the 1990s after the find of the feathered "Liaoning theropods". Archaeopteryx has had a somewhat unique role among stock prehistoric animals: just like the Dodo is the icon of extinction, Archaeopteryx has been that of evolution. Within the long-lasting debate between scientists and creationists, the latter went so far to claim Archaeopteryx fossils are just fake. According to modern knowledge Archaeopteryx is just another feathered theropod — possibly a bird, possibly a deinonychosaur, possibly more primitive than either. Highlighting its theropodian nature, one archaeopterygid skeleton with no signs of feathers was long classified in another theropod genus, Compsognathus (see later). The size of a chicken, Archaeopteryx had a long bony tail (modern birds always have stubby tails), three claws on its forelimbs, running feet with an enlarged second toe claw (this was discovered only in the 2000s), jaws with small, pointed teeth, and feathers. The main difference is that its feathers aren't just skin-covering down; it has flight feathers of very modern-looking shape on its wings and tail. It probably could glide but it is unlikely that it could flap its wings for powered flight — it didn't have the modern birds' keeled breastbone for powerful wing muscle attachment, but the usual "ventral ribs" seen in non-bird theropods. Maybe it simply used its claws to climb up trees and then glided to the next tree. Its diet probably consisted only of insects and small vertebrates. Its classic status as "the first bird" is merely traditional at this point, and the start of the "bird lineage" within the theropod branch depends on the chosen criteria to define what’s a bird and its exact position. Still, it remains one of the most ancient known dinosaurs found with imprints of feathers. In media, Archaeopteryx is well-established as the "first bird". It will fly like a bird and perch like a bird, neither of which was possible for the real-life Archaeopteryx. Media archeopteryges will lack the sickle claws on their feet, and possibly also their wing-fingers and teeth. Expect also to see them with naked heads, making them resemble feathered lizards. Actually, their heads would have been almost totally feathered like deinonychosaurs and most modern birds (see Raptor Attack).
- Entry Time: 1861
- Trope Maker: Darwin's On the Origin of Species
Lighter and softer: Ornithomimus, Struthiomimus & Gallimimus, aka the "Ostrich-dinosaurs" **
Ornithomimus was a small, agile animal (the antithesis to the classical Mighty Glacier dinosaur) that lived in North America between 75-65 million years ago, in the Late Cretaceous. It was 12 feet/3.5 meters long and weighed around 220-330 lbs/100-150 kg. Its shape was similar to a long-tailed ostrich. It had a long neck with a birdlike skull and a toothless beak. The brain and eyes were large, possibly an adaptation to support quick movement. Its tail was very long, balancing the animal when running. The legs were similar to modern running birds, with short muscular femurs, elongated tibias/shins, and three toes each. With this anatomy it may have been the fastest non-avian dinosaur, easily capable escaping the bigger and clumsier Tyrannosaurus rex. Ultimately confirmed to be feathered, which was already suspected for some time.note Ornithomimus ("bird-mimic") is the prototype of the Ornithomimids, often nicknamed "Ostrich-dinosaurs". This family also includes Struthiomimus and Gallimimus among the others. The former ("ostrich-mimic") was almost identical to the namesake of the family (it had only longer forelimbs), was just as fast and quick, and lived in North America in the same period (the two were actually considered the same animal at the time ornithomimids first entered pop-media, in 1940). They were definitively recognized as distinct only in the 1970s. The genus Dromiceiomimus ("emu-mimic") was briefly separated from Ornithomimus in the same decade, but has since been merged back in the 2000s. As their names suggest, Ornithomimus and Struthiomimus have been recognized as birdlike dinosaurs since their very first discovery. Ornithomimus was the first described (USA, 1890, during the Bone Wars), while Struthiomimus was named later (Alberta, 1917). The first U.S. remains were fragmentary, though, and complete ornithomimids came to light only in the early 20th century. They were described as fast-moving and graceful even before the Dinosaur Renaissance, and usually portrayed with erect tail and horizontal body, unlike the giant bipedal dinos with their upright bodies and dragging tails. Unusually for dinosaurs, media have never shown ornithomimids as scary killers dangerous to humans. This can be due to their lack of teeth that make them look harmless. Their actual diet is still uncertain, as no stomach remains are known for now. Their large numbers, among other things, seems to indicate that they were mainly herbivorous with insects, eggs, and small animals as a supplement. In popular media they have often been depicted as plant-eaters and/or insect-eaters (very rarely as meat-eaters). But the most common pop-portrayal has shown them as egg-stealers outwitting larger dinosaurs, like in The Land Before Time (possibly due to confusion with Oviraptor). Even though they might have eaten some eggs, there is no evidence this was a major part of their diet. Furthermore, not being maniraptorans ("robbing hands"), they probably couldn't grasp things so easily as the latter did. Today, the "robbing" role is more often attributed to the true maniraptoran oviraptorosaurs (see below). Finally, in the early 2000s it was suggested ornithomimids were filter-feeders like flamingos (as seen in Prehistoric Park), but now this hypothesis is disproved. Discovered in the 1970s, Gallimimus ("rooster-mimic") was one of the largest ornithomimids — 20 ft/6 m long, with some reports of sizes up to 8 meters long, as long as several giant predatory theropods. It lived in Mongolia in the late Cretaceous, 70 mya. Apart from its longer, blunter snout and slightly shorter legs, its appearance was that of an enlarged Ornithomimus. With its large fossil record Gallimimus has become a common sight in dino-books since the '80s, but entered the pop-consciousness only after Jurassic Park — the film, not the novel, which has the hadrosaur Maiasaura in the stampede scene. Like Velociraptor and Dilophosaurus, Gallimimus became one of the Stock Dinosaurs immediately after the movie, but has replaced Ornithomimus and Struthiomimus in pop-culture only partially. Fictional works from after 1993 can still add Ornitho- or Struthio- (or even "Dromiceio-") to their -mimuses, instead of Galli-.
- Entry Time: 1940 (Ornithomimus/Struthiomimus); 1993 (Gallimimus)
- Trope Maker: Fantasia (Ornithomimus/Struthiomimus); Jurassic Park film (Gallimimus)
Cannibal, or hero? Coelophysis *
One of the first true dinosaurs to appear on Earth, Coelophysis lived in Late Triassic North America 216-203 million years ago, although fragmentary material suggests a near worldwide distribution lasting up to 188 mya. Described during the Bone Wars from some pieces of bone, today it is by far the most abundant early theropod in the fossil record. In the 1940s, a whole graveyard with hundreds of specimens was found in New Mexico; they're widely theorized to have died all together in a flood, though this is not certain. Coelophysis was a slim, fast-running dinosaur growing up to 10 ft/3 m, and weighing about 30 kg. Coelophysis looks like a fragile animal, with a narrow head, weak jaws with small pointed teeth, a long, stork-like neck (sometimes incorrectly described as "snake-like"), and an elongated, thin body. As an early theropod, Coelophysis was not very closely related to birds. For example, it had still a remnant of the fourth digit on each hand, and the presence of feathers is uncertain. It it had them, they were surely "proto-feathers" or down-like structures, not modern-looking feathers. note Still, it had bird-like features showing how far back in time the dinosaur-bird link goes. Its skull and hindlegs were similar to the more evolved theropods; its bones were hollow and had airsacs within them (its name just means "hollow frame"); and it even had a wishbone, a typically avian trait. Coelophysis probably hunted down small prey, which it swallowed whole: lizards, dinosaur nestlings, fish, insects, proto-mammals, and whatnot. In the Triassic the top predator role was played by non-dinosaurian archosaurs like Postosuchus or Rutiodon, or larger theropods like Gojirasaurus. Coelophysis is often described as a "successful underdog" which finally managed to outcompete non-dino archosaurs, anticipating the following domination of the dinosaurs in the Jurassic world. It is usually portrayed as a gregarious animal that lived and sometimes hunted in packs; although pack behavior is possible, pack-hunting is unlikely. In many works it's said adult Coelophysises ate younger members of their own kind (or even their own young) during famines. This because bones found in the stomachs of adult specimens from the aforementioned "graveyard" were reported to belong to young Coelophysis, leading to the dinosaur being described as cannibalistic. Later studies have determined that the bones in question were animals of other species. This doesn't mean that Coelophysis didn't occasionally eat their own kind (crocodiles and eagle-nestlings do this after all); only that there isn't any fossil proof anymore. Together with the large herbivorous Plateosaurus, Coelophysis is the dinosaur you're most likely to see in those documentary works portraying the Triassic Period, to show how the earliest dinosaurs looked (even though in Real Life there were many other dinos in the Triassic, some even more primitive: Staurikosaurus and Thecodontosaurus are two examples). In these works, the smaller Coelophysis is used to represent the very start of the dinosaur evolution, while the bigger Plateosaurus represents a more advanced/enlarged stage. An excellent example of all this is the first episode of the TV documentary Walking with Dinosaurs, in which the two animals are shown living together in North America (in Real Life only the coelophysis was North American, the plateosaur was found in Europe). On the other hand, Coelophysis has been far less common in fiction or other more popular media, since it is too humble-looking and generic to be interesting; the best-known appearance may be "Spot" from the 1974 children's television series Land of the Lost.
Dinosaurs as pets: Compsognathus **
Compsognathus was native to Europe 150 million years ago and lived alongside the famed "first bird" Archaeopteryx, in the same habitat made of small islands. Like the "Urvogel", it was one of the first dinosaurs described, in the same year as the latter. The first Compsognathus specimen was found in Germany in the Solnhofen site (like Archaeopteryx). The second known specimen was found later in Southern France. Both skeletons are nearly-complete and still-articulated (again like Archaeopteryx), but haven't preserved imprints of skin. Though little-known, Compsognathus may have been the first non-bird dinosaur ever found from an almost-complete skeleton. Only 4 ft long and weighing few kilograms, Compsognathus is the smallest Stock Dinosaur, not counting Archaeopteryx (and Microraptor, which can be considered a real stock at this point). Like Coelophysis above, its frame was that of a generic small theropod with no sickle-claws on its feet. Compsognathus was more evolved and bird-like than Coelophysis however, with a shorter head and neck, a more compact body, and only three fingers on each hand. Old portraits in popular dino-books have typically depicted the "compy" with two-fingered T. rex-like hands, and sometimes even with fin-like hands: these errors were due to the incompleteness of the two known specimen's hands. If you hear about it in documentary media, it will likely be for two things: its former record of "the smallest dinosaur" (classically described as chicken-sized because its first-found skeleton was only 2 ft long, but was from a subadult), and its former status as "the closest relative of Archaeopteryx" (despite similarities, it was possibly less close to birds than tyrannosaurs). Another compsognathid, Sinosauropteryx from Early Cretaceous China, shows downy covering around its skeleton but not pennaceous feathers; this was probably the same for Compsognathus, too. Compsognathus has always been a regular sight in dino-books, in which is usually shown as a solitary hunter of insects, small vertebrates, and sometimes adult archaeopteryges (that last depiction is unlikely though).note Even though its tiny size doesn't make the "compy" particularly menacing for humans, popular media have equally managed to transform it into a "miniature terror". In 1997, The Lost World: Jurassic Park made it a deceptively cute critter which attacks in huge packs and kills humans with a paralytic bite. While Compsognathus may have been cute (its name means "dainty-jaw"), in Real Life there is no indication of social behavior, and its jaws and teeth were strictly adapted to catch small prey. The fact that the original specimen's stomach cavity contained only a small lizard would tend to support this.
Oviraptor lived in Late Cretaceous Asia 75 million years ago. This dinosaur is another demonstration about how Science Marches On is a crucial factor in popular dino potrayals. First discovered in Mongolia in 1924 together with Protoceratops and Velociraptor, this toothless theropod was originally thought to be an ornithomimid. Since the 1970s it is classified into its own family, even more closely-related to birds. Oviraptor was distinct from ornithomimids by having a rounder, stockier beak, a shorter tail, and the small forth reversed toe on each foot present in most theropods (birds included), which was lost in ornithomimids. Unlike the latter, it was a "maniraptoran" theropod: its forelimbs were bird-like, and it's considered to have had a very bird-like covering of feathers, with feathered wings and a feathered tail fan. It was given the name Oviraptor, meaning "egg-thief," because the first specimen (a crushed skull) was found next to a clutch of eggs which were thought to belong to the small ceratopsian Protoceratops. In the 1990s the eggs were found to contain oviraptor chicks: the specimen was brooding its eggs. This was further confirmed few years later, when an oviraptorid skeleton was found just above a nest full of the same kind of eggs. Before the middle 1990s scientists used to describe Oviraptors as a specialist nest-robbers, and documentary works have typically shown them frequenting the nesting ground of herbivorous dinosaurs (especially those of Protoceratops), grasping the eggs with their prehensile hands, and fleeing away from the angry mothers when they're discovered. As said above, the "robbing" behavior has often been attributed to ornithomimids as well, but Oviraptor was considered specifically adapted just to eat eggs and no other kind of food, with its robust bill that was thought a specialization for breaking eggshells. Today, the actual diet of oviraptorids is a matter of speculation — they might have been fruit-eaters, predators, or both; they might even have fed on eggs if given the chance. Since the first discovery the oviraptor has also notably changed its look: paleo-artistic works from before the 1990s usually depicted it featherless (like every other non-bird theropod), and with a strange protruding palate with a pair of teeth at the tip. In works made in this period you can see the animal with two different head-shapes: the one with a small bony bump on its nose (inaccurate), and the most familiar today, with a square bony flat crest. (see also Prehistoric Life). However, since the 2000s the most complete Oviraptor skeletons have been reclassified in a brand new genus, Citipati: the familiar square-crested image probably belongs to this new genus. Note that this is not a "Brontosaurus"/Apatosaurus case: the genus Oviraptor is still valid, it's just that many specimens that used to be considered to belong to the genus are now considered Citipati, and that the actual Oviraptor is very unlike its depictions, lacking any known crest and being far smaller that its emu-sized relative. Unlike ornithomimids, oviraptorids have attracted the attention of dino-writers only since the 2000s: after the appearance of the feathered, non-egg-stealing scientific depiction. And yet, expect to see them portrayed in the older inaccurate way nonetheless (and you can also see oviraptorids and ornithomimids mixed up with each other). E.g. in the 2000 Disney movie Dinosaur, a featherless "Oviraptor" steals Aladar's egg, but loses it before it has a chance to eat the contents. More recently, an "Oviraptor" appeared in The Land Before Time TV series: feathered and not egg-stealing. Both portrayals show the oviraptors with a Citipati-like tall crest on their head. Dinotopia lampshades the animal's Science Marches On story showing it in two variations: the featherless "Oviraptor" and the feathered "Ovinutrix" ("egg-nurse").
The big brain: Troodon and the "Dinosauroid" *
This has been another complex case of Science Marches On as well, but also an astounding example about how imagination can be a very influential factor even for people studying dinosaurs seriously. Living in North America some 75-65 million years ago alongside giant tyrannosaurs, Troodon was a small dinosaur only around 7.9 ft/2.4 m in length and weighing some 110 lb/50 kg. It was still the largest member of the troodontid family, a sibling family to Dromaeosauridae and among the closest relatives to birds. Troodontids looked a lot like dromaeosaurids, including being covered with feathers. Specifically, Troodon was rather similar in shape to the Real Life Velociraptor; you can tell apart the two by observing the troodont's shorter head, smaller teeth, eyes pointing forwards note , shorter tail, longer legs, and less formidable sickle-claws on their second toe. Since the 1980s Troodon has attracted scientists' attention because it shows several very specialized anatomical traits: it had larger eyes and ears than most dinosaurs, perhaps indicating nocturnal habits, and its brain was relatively large for a non-bird dinosaur as well. Its forwards-pointing eyes show binocular vision similar to modern owls; many old portraits showed it with bulbous eyes with cat-like or even gecko-like pupils, almost resembling humanoid aliens (remember that last detail, we'll return to it at the end). More realistically, it had bird-like eyes with round pupils. It used also to be imagined with opposable thumbs making its hands like an eagle's foot to better grasp its prey, but this is controversial. The troodont's actual diet is still debated: with its small, relatively blunt teeth, it was likely a mostly-carnivorous omnivore, though it used to be portrayed as a specialist small-prey hunter (mammals, dinosaur nestlings). Some have even hypothized it was an herbivore, but this is now mostly discarded. At the other extreme, some depictions attribute to it dromaeosaurian traits such as pack behavior or the ability to kill large prey, which weren't possible in Real Life. Troodon 's toothed jaws and sickle-claws were too weak to tear the meat of large living animals. The first find was a single tooth (hence the name Troodon, "wounding tooth"), one of the very first North American dinosaurs found (1856), but it was initially believed a lizard, then a pachycephalosaurian. Meanwhile, a small theropod, Stenonychosaurus, was described in the 1920s, and classified as a generic "coelurosaur". In 1987, paleontologist Phil Currie found the two animals to be one and the same, and Stenonychosaurus fell in disuse in favor of Troodon. If you read popular dino-books written before the 1990s, you'll probably find the name "Stenonychosaurus" more often. note Also in the '80s, scientists found the troodont’s brain to be the biggest for its body size among all dinosaurs: this gave it the reputation of "the smartest dinosaur" in popular books note — despite this, its brain was still smaller than most modern birds. Despite an emergent tend to show up in documentaries, the troodont’s presence in fiction has been only occasional, and not related to the actual animal but to that could be called its "altmode". In 1982 note , Canadian paleontologist Dale Russell conjectured a possible way that descendants of Troodon could have evolved had it not gone extinct along with the rest of the dinosaurs. If its brain had kept increasing in size, today it would have been comparable to a human's. Combined with further evolution of its bipedal movement, binocular vision, and semi-manipulative hands, the resulting "Dinosauroid" was proposed to be a blend of (featherless) dinosaurian and humanoid features. The Dinosauroid has made a few appearances in novels and TV series; its Real Life dino-ancestor usually gets mentioned. It's worth noting that the Dinosauroid model resembles the Sleestaks of Land of the Lost (1974-1977), possibly a case of Ascended Fanon. Recently, Troodon's validity has been called into question, as the type specimen only consists of teeth. While Troodon falls into taxonomic limbo, many fossils assigned to it have been moved into genera of their own, while Stenonychosaurus has been resurrected, much as Brontosaurus was.
- Entry Time: Uncertain
- Trope Maker: the "Dinosauroid" hypothesis in the 80s.
Four-winged dinosaur: Microraptor *Here's one of the latest entries in the Stock Dinosaurs realm. Discovered in year 2000, Microraptor is one of the "Liaoning coelurosaurs" (see Prehistoric Life - Birdlike Theropods), named “small thief”; as the "-raptor" suffix suggests, it was a dromaeosaurid. It was a find that surprised not only casual paleo-fans but also the entire paleontology community. And not because it was a feathered dino fossil (such animals were already known from the same site), nor just because it was the smallest non-avian dinosaur known at that point (merely 1.5ft long, but this record is contended now by other non-avian maniraptors). It was its unique body-plan that astonished us all: a four-winged dinosaur! More precisely, its hindlimbs had a feather covering incredibly similar to that of its forelimbs, giving it its unbelievable appearance. These wings had the same structure as the wings of true birds, with asymmetrical, vane-like feathers on the forelimbs, likewise on the hindlimbs, and placed in a "fan" at the tip of its long tail: in short, very similar to the kind of plumage of the well-known Archaeopteryx (itself recently found to have had remnants of such large feathers on its legs). Of course, paleontologists and dino-fans have begun Wild Mass Guessing about its way of life. Since its discovery, Microraptor has been suggested to have been a tree-climber, with forelimbs as developed as the hindlimbs, both fitted with robust claws apt for climbing upright tree trunks; however, a study published in 2011 suggests it might have been terrestrial instead. The way it traversed the air is also controversial; with true flight like modern birds, or just simple gliding like modern “flying” squirrels, “flying” fish and “flying” lizards? Currently many scientists think Microraptor was actually a flier (although not as good as modern birds): not only that, it seemed to be even better adapted for flight than Archaeopteryx. If this is true, it would mean that flight evolved before the appearance of the so-called “first bird”, because Microraptor was less close to modern birds than Archaeopteryx was. And since flight was achieved in basal dromaeosaurids, this would mean that... yes, Velociraptor and all other dromaeosaurids may have descended from flying ancestors! One scientist did go Up to Eleven declaring that all maniraptorans descended from flying ancestors: this would mean, Troodon, Oviraptor, and even the huge Therizinosaurus were ancestrally creatures of the air, which, like ostriches or rheas, returned to a more ground-level way of life and increased their size. Whatever the case was in Real Life, Microraptor immediately became the center of much interest soon after the year 2000, rapidly becoming popular in illustrated books (also because it was the considered the smallest dinosaur at the time); it became even more widely-known after being included as one of the main animal characters in the aforementioned Prehistoric Park (where it was portrayed with the classic, splayed-limbs gliding style, now known to be anatomically impossible). Soon afterwards, it started to gain attention in the broader pop culture world, and it could at this point be qualified as a true Stock Dinosaur (even if only in the Rarely-Seen section, with one asterisk on the top).
Wolverine-clawed dinosaur: Therizinosaurus *Therizinosaurus could be considered the non-identical twin of Deinocheirus: it was a colossal yet awfully bird-like theropod, just like Deinocheirus; specialized to a non-big-prey-based diet, just like Deinocheirus; was discovered in Late Cretaceous rocks from Mongolia, just like Deinocheirus; is known mainly from forelimbs and few other bits, just like Deinocheirus before 2014; entered the dinosaur list around the same time as Deinocheirus; and, last but not least, it is another candidate for the title of biggest theropod, just like Deinocheirus! But, unlike Deinocheirus, Therizinosaurus was not a giant ornithomimosaur — but still an equally huge freak. Discovered in the 1950s but not recognized as a dinosaur until the 1970s, its forelimbs were slightly shorter but more powerful than those of the giant ornithomimosaur. But Therizinosaurus had an additional curiosity, one that made it even more awesome: three scythe-like claws on each hand (hence its name, "scythe lizard"), some as long as a human arm. In short, it had the biggest nails known so far within the entire Animal Kingdom. One of these oversized claws was in fact the first known find, and for several years, scientists thought it belonged to a giant marine turtle. With such powerful weapons, Therizinosaurus has in the past received the same treatment as Deinocheirus. Some old drawings went as far as to show our "scythe-dino" as a giant carnosaur or deinonychosaur with sickle-claws on each foot (if Therizinosaurus was really shaped that way, it would really have been the most badass dinosaur one can imagine...). More accurate analyses made at the beginning of the 1990s definitively debunked these fantasies: we now know with a good level of certainty that Therizinosaurus was a bulky-bodied, round-bellied, and quite slow-moving animal that used its claws mainly to pull down branches. Furthermore, its jaws were arguably weak with a rounded horny tip and small grinding teeth similar to those seen in its relative Segnosaurus. This obviously doesn't lessen its general coolness: even with this new shape, Therizinosaurus remains an odd-looking, powerful beast, and thanks its massive body, it might even be the biggest and heaviest theropod ever discovered, weighing even more than the famous Spinosaurus. As we've long done with Deinocheirus, we dino-fans are patiently waiting for exciting new remains of our "Wolverine Claws-osaurus" to be excavated. Meanwhile, a special spinoff of Walking with Dinosaurs from 2002 temporarily recreated our imagination in CGI: in the episode titled "The Giant Claw" Nigel Marven talks about Therizinosaurus, lampshading its whole Science Marches On story from a mighty carnivore to a Gentle Giant. Nigel is in Late Cretaceous Mongolia searching for the possessor of the eponymous "giant claw", which the zoologist believes to have pertained to a fearsome predator. After several adventures with other famous dinosaurs of the habitat (Protoceratops, Velociraptor, etc.), Nigel witnesses a fight between Therizinosaurus and Tarbosaurus: even though the former unexpectedly reveals itself to be an herbivore, it easily defeats the tyrannosaur by slapping it in the face with its scythe-claws, obliging the predator to flee. Finally, the therizinosaur licks Nigel’s face. Really!
- Entry Time: 2002
- Trope Maker: Chased by Dinosaurs
Other small or birdlike theropodsSorry, these ones aren't here. If you're looking for Dromaeosaurus, Ornitholestes, Mononykus, Deinocheirus, and others, see here. Or here.
As easy as it is to forget, birds are dinosaurs as well. Specifically, birds are members of the maniraptoran clade, which also includes the aforementioned dromaeosaurs, troodontids and oviraptorans. Yes, that means that dinosaurs are technically not extinct! These contain the only Cenozoic dinosaurs on this list.
Monster penguin and toothy seagull: Hesperornis and Ichthyornis*Hesperornis and Ichthyornis are two of the most famous (and the two most commonly depicted in books) Mesozoic birds (not counting Archaeopteryx, of course), both from Late Cretaceous North America. Since Hesperornis is far cooler, here we'll mention it first. Hesperornis lived in the same habitat in which pteranodonts, mosasaurs, elasmosaurs and Archelons roamed: the shallow inland sea which used to cover the American Midwest at that time, dividing North America in two parallel strips of land running from the Arctic down to the south. Despite living in an early period, Hesperornis was already a very derived bird. 6 ft long (the size of a human), it was flightless, with vestigial wings, short splayed legs for swimming, a long neck, and a long beak with small true teeth. It spent most of its life in water, but returned on land to lay its eggs. Once, the hesperorn was shown as a sort of proto-penguin with an erect pose; we know now its legs were too weak to support its body like this, and the animal is now portrayed as more similar to modern grebes and loons. It was once thought to have had palmated feet like a loon, but it's more likely it had flattened toes like a grebe. Similar to it but smaller, Baptornis also lived in the same shallow seas. The much smaller, far less striking Ichthyornis lived in the same age and habitat of the former, but this time we're faced with a sorta toothed, long-billed proto-seagull. The ichthyorn's lifestyle was arguably similar to modern flying sea birds, catching fishes in flight or maybe diving under the sea to pursue them like modern boobies. You can almost be certain that if pterosaurs are involved, these little guys will be depicted as pests who like to steal food from them, a behavior that while possible, has never been proven. Both ichthyos and hesperos sometimes fell prey to large marine reptiles, as shown by remains of stomach contents. Ichthyornis and Hesperornis were (almost) full birdies at that point, and if alive today, they'll be taken for components of modern avifauna. In some artistic works, both Hesperornis and Ichthyornis are depicted as black and white like modern gulls or penguins, but their real coloration is totally unknown. While these two do feature prominently in dinosaur books and sometimes documentaries (Hesperornis, for example, was the token prey animal in Sea Monsters and Ichthyornis got a bit part in Dinosaur Planet as a scummy scavenger), their presence is rare in more mainstream media, presumably because, besides their teeth and Hesperornis' large size, they don't have a lot of cool points. That hasn't stopped folks from trying (for better or worse). An Ichthyornis, creatively and perhaps fittingly named "Ichy" appears as a one-shot villain in the fourth Land Before Time movie, accompanied by an equally villainous Deinosuchus. Ichthyornis also cameos in Dinosaur, erroneously depicted as duck-like creatures rather than seagull-like, and are among the many factors contributing to the start of the movie when they attack the mother Pteranodon carrying Aladar's egg, causing her to drop it. In Primeval, Hesperornis appears as an aggressive but non-malicious creature that kills a plumber after its anomaly appears in someone's flooded basement. It's portrayal there is probably one of the worst of any prehistoric bird — to the point that the creature designers had it featherless and standing upright.
- Entry time: 1872 (both)
- Trope makers: The Bone Wars (both)
Dinosaurs continued to rule: Gastornis (once called "Diatryma"), Phorusrhacos, & Titanis, aka the “Terror Birds” *Long-standing paleo-fans will remember the name "Diatryma" for sure: that large, flightless, large-headed predatory bird who used to hunt the small "horse" Eohippus in so many paleo-artistic depictions. Well, now poor "Diatryma" seems having definitively disappeared... but luckily, it hasn't, as such: it's simply changed identity. Now we have to call it Gastornis (a far less awesome name, we've got to admit, but... never mind.) Whatever name should be used, this is actually one of the most enigmatic extinct birds. Recently, it was shown that it wasn't even carnivorous at all: its strong beak wasn't hooked like an eagle's, and its body frame was stocky, seemingly slow-moving. It only was an herbivore that used its bill to crack nuts and cut vegetation — making erroneous the Bizarro World portrayal in Walking with Dinosaurs where birds were shown ruling mammals. However, this does not mean it was the Gentle Giant news articles claimed it was. Ostriches and cassowaries are herbivores too, but they're also some of the few birds that have been known to kill people. And Gastornis not only grew to their size, but it also had a powerful beak that would've been useful for fighting off the land-dwelling crocodiles that were the true dominant predator. Needless to say, the gastorn/diatryma was a literal and figurative giant in its forested world, 40 million years ago: while mammals were still small, some birds grew to large size. With the Phorusrhacids, on the other hand, we have no doubts this time: thanks to their light weight and slender running legs, they were active hunters of small mammals. Not only that, with their strongly hooked, very eagle-like bills, they did not swallow their prey whole. They were once thought to have one clawed fingers protruding from each of their tiny wings note , for uncertain purpose. However, it was later discovered that their living relatives, the seriemas (see below) have similarly-shaped wings and lack wing claws, making these fingers unlikely. Perhaps the most amazing-looking among all prehistoric birds, they have recently been nicknamed terror birds in pop-documentaries (for example, Prehistoric Park). Originally from South America, they have left a legacy in our modern world as well: the closely-related seriema is a medium-sized South American bird whose shape and habits resemble a miniaturized "terror bird". The prototypical South American Phorusrhacos (often misspelled "Phororhacos") and the North American Titanis (which first originated in South America as well) are the two stock species of the family. Despite being only distant relatives, and the aforementioned Science Marches On, expect Gastornis to be lumped in or confused with the true "terror birds" anyway. In fact, Gastornis was actually more closely related to waterfowl than to the Phorusrhacids. The most notable appearance in non-documentary fiction is in 10,000 BC, where they fill the role of raptors. Never mind that, even if species like Titanis did live alongside humans, they never made it to Egypt, where the film appears to take place.
- Entry Time: 2001
- Trope Maker: Walking With Beasts
Symbol of Man's Folly: The Dodo (Raphus cucullatus)***Perhaps the most recently extinct stock dinosaur (in a technical sense, anyway), having been wiped out before 1700, the dodo was a turkey-sized flightless pigeon that lived on the isolated island of Mauritius. The pigeons it was descended from almost certainly flew to the island, but because there were no land predators, the dodo evolved secondary flightlessness, no longer needing the luxury of flight—which may have been an evolutionary mistake, considering what happened when humans and their pets showed up. Its diet likely comprised of fruit and seeds, as well as possibly insects. The circumstances of the dodo's extinction are commonly misinterpreted. The most common explanation was that they were simply unable to comprehend that humans were NOT friendly and thus were hunted for their meat until they all died out. In reality, while it is likely that these birds were a little too brave for their own good (many modern birds on Mauritius still can't take the hint), they certainly weren't helpless creatures—their beaks were very large and sharp and reportedly were capable of delivering painful nips if the dodos got agitated. Additionally, the meat of the dodo was said to have been very tough and oily, making it very unpalatable for human consumption. It is now thought that the demise of the dodo came along because of the invasive species brought into the island—rats and cats ate the babies and and eggs while pigs and dogs ate the adults. In addition to the previously mentioned flightlessness and possible Fearless Fool status, dodos were very slow breeders who couldn't repopulate faster than invasive predators were eating their young, so it wasn't long before they all disappeared. Dodos are frequently thought of as having been stupid creatures, to the point that we have a trope for this portrayal and a proposed origin for its name was a Dutch word meaning "simpleton". We now know that dodos were no more or less intelligent than any other pigeon and that because it was a pigeon, a more likely origin for its name was the sound it made, which most likely sounded like a soft "doo-doo". The dodo had a cousin on a neighboring island called the Rodriguez solitaire—which sadly also went extinct—and its closest living relative is the Nicobar pigeon. Other extinct birds from these islands included a giant ground-dwelling parrot, a harrier, a sparrowhawk-like owl, and a flightless ibis. The dodo is probably the most famous extinct animal that didn't come from the Mesozoic or the Pleistocene Ice Age, to the point that a notable euphemism for death or obsolescence is "going the way of the dodo" or "dead as a dodo". On a more serious note, the dodo's extinction is one of the things that has motivated humankind to try and be more environmentally minded—after all, we were able to wipe this innocent, goofy-looking bird out of extinction, who's to say we won't end up wiping out more species? Sadly, not everyone got the hint, not to mention we lost many species before the dodo too. The dodo's fame probably started with Alice's Adventures in Wonderland, when an anthropomorphic dodo appears among the animals Alice encounters in Wonderland. The Dodo in the book is said to be an Author Avatar for Lewis Carroll. Dodos have appeared all over the media, usually depicted in the outdated, fat, stupid waddling birds portrayal, as opposed to the thinner and more bold birds that we know they were now. They'll often be put into a post-dinosaur world for comic relief—for example, in Ice Age where the dodos are depicted as possessing almost suicidal stupidity. Most media will also forget that the dodo is a pigeon. Regardless, the dodo will almost never be depicted as a Prehistoric Monster, unless it's mutated or something, and even then it will be seen as a joke rather than a threat, as was the case in Series/Primeval.
- Entry Time: 1865
- Trope Maker: Alice's Adventures in Wonderland
Other birdsSorry, these aren't here. If you're looking for Aurornis, Confuciusornis, Teratornis, Argentavis, Osteodontornis, and others, see here.
Long-necked plant-eatersThe sauropodomorphs ("sauropod-shaped") include dinosaurs with long necks. Most of them pertain to the sauropod subgroup, while all the other members are traditionally called prosauropods.
Sauropods are the (mostly) gigantic quadrupedal plant-eating dinosaurs with long necks and tails. Some of them were the largest land animals that ever lived, but not quite as massive as they seem: the weight was brought down significantly by a system of air sacs in hollow bones, similar to theropod (and hence, bird) skeletons. Since sauropods are rather similar to each other in size and appearance, only a few of them will usually be identified/identifiable in Fictionland: "Brontosaurus", Brachiosaurus, and Diplodocus. Their iconic status is due to several factors: the early time of their discovery, the relative completeness of their remains, and for having held some "record" in the past — Diplodocus has long been the "longest" land animal ever, the brachiosaur the "tallest" and "heaviest", while "Brontosaurus", being the very first mounted sauropod in a museum, was the first that became popular. Today scientists recognize more than one hundred sauropod genera, but pop culture ignores most of them. All stock sauropods are from Late Jurassic North America (even though the best-known brachiosaur species was found in Africa), but in Real Life sauropods lived worldwide from the Early Jurassic up to the end of the Dinosaur Age. However, in recent years some other sauropods have received attention in media, but only because were said to be "Longer! Taller! Heavier!" than the traditional record-holders. Some really were, but other weren't even real animals. You can find them here classified as "little stock".
Common inaccuracies in sauropod portraitsIf a writer relies on pre-Renaissance science, a featured sauropod is up to its armpits in water and living in swamps while lazily munching some swamp weeds. Before The '70s, the dominant but incorrect hypothesis said they needed to spend most of their time in water to support their massive bulk and to escape the (allegedly) non-swimming theropods. Modern science says sauropods were terrestrial (though able to swim in an elephant style if needed); note if they were really swamp-specialists, they would have had a high chance of slipping in the mud with fatal consequences. If lucky enough to survive the fall, they'd starve to death from lack of nutritious food. note If submerged, their ribcages wouldn't even be capable of expanding due to water pressure, suffocating them. And the alleged weakness of their teeth and the high placement of their nasal openings are not real proof for an amphibious, swamp weed-eating lifestyle.note Finally, footprints show carnivorous dinosaurs were capable of swimming using their hindlegs (as shown by the aforementioned alleged Dilophosaurus footprints), making the sauropods' fleeing in water to escape them potentially useless. Another common mistake when portraying sauropods is to show them with elephant-like nails or hooves, falling straight in Most Writers Are Mammals. Actually, sauropods had true claws. They usually had a thumb-claw on each forefoot (which was narrower than in modern elephants) and three claws on each hindfoot (which was broader and more elephant-like than the forefoot). Even so, in most portrayals that do show clawed sauropods, they usually have four or five claws on each foot. More related to Reptiles Are Abhorrent is the tendency of depicting sauropods' necks as serpentine: you'll even find brontosaurs using them like snakes when attacking their prey. Actually, their necks had relatively few vertebrae, like a giraffe's, and were relatively stiff (expecially if compared with their flexible tails, which often had 50-70 or more bones).
Deceptive dinosaur: Apatosaurus, aka "Brontosaurus" ***Lived in the Late Jurassic (154 to 150 million years ago) in what is now the USA. It was large, 75 ft/23 m long and weighing at least 23 metric tons (equal to roughly four elephants). The neck made up about a quarter of its total length, the tail about half. Apatosaurus’ overall size is often exaggerated in popular writing, for instance by claiming that it weighed as much as 10 elephants, or even that it was the largest dinosaur (one could assume the writer knew only one type of sauropod). The head was small and slender; the teeth were peg-like and found only at the tips of the jaws. The neck had more than 10 vertebrae, and was of average length but wider than in other sauropods. The body was stocky and deep; the hips were taller than the shoulders. The legs were robust (even more so than in most sauropods), the hindlimbs longer than the forelimbs. The tail was very long (about 80 vertebrae), thin and whiplike near the end. Apatosaurus is one of the few sauropod genera already known to science when dinosaurs entered pop culture in the 1900s. Marsh described the first species in 1877, in full Bone Wars context, but its first remains were incomplete. Just two years later Marsh described a second Apatosaurus species as a distinct animal, "Brontosaurus". It was the latter that became the iconic image of a sauropod for most the 1900s. Apatosaurus and Tyrannosaurus were the ultimate stock dinosaurs in their respective roles as herbivore and carnivore, and contenders for the title of THE overall most iconic dinosaur. In classic dino-stories the "brontosaur"'s designated role is the Gentle Giant (while T. rex is the Big Bad and Triceratops the badass or The Hero)... unless Everything is Trying to Kill You, of course. In the classic version of King Kong, brontosaurs are meat eaters. Why is Apatosaurus the correct name for this dinosaur? Because scientific rules say if one animal is assigned two scientific names, only the first one is valid, and "Apatosaurus" was coined two years before "Brontosaurus". The latter means "thunder lizard" (probably a reference to the booming sounds sauropods might have made when walking), while the former means "deceptive lizard" — a much less cool name but, surpisingly, the most apt one. And then, why has just "Brontosaurus" been the traditionally most popular sauropod, despite being smaller than Brachiosaurus and shorter than Diplodocus? Probably because the first-ever mounted display of a sauropod skeleton (erected at the American Museum of Natural History in 1905) was based on a mostly complete Apatosaurus skeleton, with missing parts borrowed from other sauropod specimens, which also gave the display skeleton a short, boxy head and blunt tail, both incorrect. However, the Museum chose to label the display "Brontosaurus". Other museums followed suit with similar "Brontosaurus" displays. Popular writing and dino-art kept spreading the incorrect name, and the ghost of "Brontosaurus" still haunts Apatosaurus, as does the image of the short, round head — even though the long-narrow-head portrayal is starting to become more known among general public. note Peter Jackson's King Kong (2005) referenced this situation by having a newly-discovered dinosaur on Kong Island be named Brontosaurus in the special features on the DVD. A nice homage, but in reality, once a name is used, even if it's invalidated, it can never be used again for a new animal, lest later researchers be left with no idea which Brontosaurus you're talking about. Even though the name "Apatosaurus" is today more known among common people than before thanks to documentary media, it could fall again in oblivion if recent proposals are confirmed in the next few years. Since the early 20th century the genus Apatosaurus has traditionally included Apatosaurus ajax (the first described species), Apatosaurus excelsus (excelsus = elevated), and Apatosaurus louisae, plus two lesser-known, recently-identified species. However, in 2015, a new study re-classified Apatosaurus excelsus and the two recently-identified species in the genus Brontosaurus after finding differences between "Apatosaurus" excelsus and the type species of Apatosaurus, A. ajax. If further studies ultimately confirm this proposal, both genus names will be validated, with the more famous species becoming Brontosaurus once again. Winsor McCay's famous 1914 Gertie the Dinosaur made "Brontosaurus" the very first dinosaur in cinema. Interestingly, she's shown as a land animal, anticipating a largely-discarded theory for more than half a century. Gertie has also the correct head shape for an apatosaur. However, since the "brontosaur" head was considered round at the time, some hypothesize she's actually meant to be a Diplodocus.
- Entry Time: 1905
- Trope Maker: Display at American Museum of Natural History (as "Brontosaurus")
The "longest": Diplodocus **
Living in western North America during the Late Jurassic Period (154-150 million years ago), Diplodocus was a neighbor of Apatosaurus. Both dinosaurs belonged to the same family, Diplodocidae, and many features of Apatosaurus (the whip-like tail, the skull shape, and the longer hindlimbs) are shared by Diplodocus. Unlike Apatosaurus, Diplodocus' portraits have always had a narrow-ended tail and the long head with a flattened snout typical of diplodocids (the Diplodocus' skull and tail-end have been known since the first discoveries). This means the two animals can be easily distinguished from each other in older media. note In more updated depictions, their overall profiles are the main key to telling Diplodocus from Apatosaurus. note The diplodocuses were longer than the apatosaurs, from about 80 ft/24 m to 115 ft/35 m, but weighed only about half as much (10-16 metric tons). The Diplodocus shape was more slender and elegant than the robust Apatosaurus, with a longer, slimmer neck; these differences allowed the two animals to live side-by-side and avoid competition by browsing different kind of vegetation. Diplodocus has classically been qualified as "the longest dinosaur", but this record is now contended by other diplodocids, like Supersaurus. All diplodocid sauropods had long tails, but Diplodocus took this to an extreme. Its tail was 14 m long, longer than a whole T. rex! note It has been speculated that the thin end of the diplodocids' tail could have been used as a whip directly against threats, or indirectly by making whip-cracking sounds. The tail vertebrae also had double beams (hence the name Diplodocus: "double beam") that may have protected the blood vessels inside the tail when the tail pressed against the ground. Diplodocid sauropods may have used their tails as a support together with their hindlimbs, lifting their forequarters to reach higher vegetation. Found during the Bone Wars like Apatosaurus, Diplodocus was introduced to the public courtesy of Andrew Carnegie some decades later. He sponsored an expedition that discovered a new Diplodocus species (which was named Diplodocus carnegii), note and had the remains mounted in his museum in Pittsburgh. He then donated replicas of it to museums all over the world; as a result, in some European nations (especially Britain) Diplodocus became the iconic sauropod, rather than "Brontosaurus" as in the USA. In recent years, the British Speculative Documentary Walking with Dinosaurs has popularized some recent theories about Diplodocus and sauropods in general: the straight, horizontal neck posture and the iguana-like spiky back. The first is due to analysis of the neck vertebrae using computer models; note the second arose from a discovery made in the 1990s of a diplo with imprints of horny spikes near its back. Both theories are now disputed: both the base and the end of the sauropods' necks were more flexible, and the animals may have been able to fold their necks and lift them like most modern long-necked animals. The spikes were dermic structures not related to the skeleton; dinosaurs being closer to birds than to lizards, the structures might have been spread over the animal's back like theropod feathers, instead of in a single line like an iguana's. We don't know if other sauropods had spikes, but spiky longnecks are now a common sight in books and art — as it seems, dino-artists have hard work getting rid of the "Dinosaurs Are Lizards" idea even today.
- Entry Time: 1905
- Trope Maker: Carnegie Museum skeleton and subsequent replicas
The "tallest": Brachiosaurus in the former sense **
This is the third member of the iconic sauropod Power Trio and lived along Apatosaurus and Diplodocus in Late Jurassic North America, 154-153 mya, but also (allegedly) in Africa in the same period. This detail was mentioned as a concrete proof about the Pangaea hypothesis, as brachiosaurs weren't able to cross oceans to migrate from one landmass to another. (The continents were still not completely separated from each other in the Jurassic.) From its first description at the start of the 20th century, Brachiosaurus was considered "The biggest land animal ever!" until real or alleged new sauropods were described starting in the 1970s (see below). Of course, works made after The '70s may still qualify the brachiosaur in this way (sadly, among them, even Walking with Dinosaurs, at least the original series). Generally thought to weigh between 30 and 50 tons, Brachiosaurus has often been oversized in popular books, so far as to triple its size up to 130 tons, which would make it heavier than any animal alive today, except for the blue whale. Brachiosaurs are visually distinct from diplodocids in several ways. First, their necks were noticeably longer than their tails, and their backs sloped backwards instead of forwards. Going into more detail, their tails had a thicker end lacking any "whip"; their necks were stronger, had more vertebrae and were held more vertically, like a giraffe; their teeth bordered most of their jaws and were chisel-like; their nasal openings were unfused, placed more forward, and were much wider than diplodocids' (the brachiosaur subgroup of sauropods, "Macronarians", just means "large nostrils").note Finally, brachiosaurs are almost the only non-bird dinosaurs with forelimbs longer than hindlimbs (Brachiosaurus means "arm lizard"). Sometimes these difference get glossed over in popular media, which may show brachiosaurs with diplodocid heads, necks, bodies, legs, and tails. In these cases, they might be recognizable as brachiosaurs only thanks to a more upright body-shape. As with other sauropods, it was associated with water in older reconstructions. To accommodate its upright shape, Brachiosaurus was often shown totally submerged in lakes, with only its head and, sometimes, only its nostrils above the water level, making its neck like a giant snorkel. Needless to say, this is quite unrealistic. In modern portrayals, Brachiosaurus has often been described as a "prehistoric giraffe" capable of browsing the highest vegetation that other sauropods were not capable of reaching — unless diplodocids were able to stand upright on their hindlegs. Brachiosaurs probably weren’t capable of that, having their center of gravity much farther forward, and their shorter tails didn't provide support (they would only bring a little bit of extra reach anyway). Moreover, as their greater size could have automatically protected adults even against the biggest predators, they didn't need to lift their forebody and use the front-legs to fight an Allosaurus, nor didn't they need to use their short tail to hit it to death (Diplodocus and Apatosaurus probably did both of these). However, in fictional (and sometimes even documentary) media, brachiosaurs show unusual athletic skills: in Jurassic Park, a brachiosaur rears up on its hindlegs to reach a tiny branch. In Disney's Dinosaur, the brachiosaur Baylene is able to remain in a fully erect position for 30 seconds to break the wall of a cave with her forefeet. Also in Jurassic Park, a brachiosaur is shown with an oversized head with fleshy lips, chewing vegetation like a cow. Sauropods didn't chew: their teeth were more suited to cutting plant material (in the case of the brachiosaur), or to raking it like a comb (like the diplodocids). Strangely, before the '90s Brachiosaurus was an unusual sight in cinema, much rarer than "Brontosaurus" and Diplodocus despite its record size. note Then, in 1993, Spielberg made a brachiosaur the solemn show-opener of his first film (Crichton's eponymous novel had Apatosaurus in this role). Thanks to this (and possibly to its cooler look), the brachiosaur has perhaps become the most frequent longneck in popular works since then, and many "young" dino-fans now consider it THE iconic sauropod in pop culture (sometimes even using the word "brachiosaur" generically as if it were a synonym of "sauropod"), while pre-JP dino-fans still tend to think of "Brontosaurus" (in the USA) or Diplodocus (in Britain) in these roles. Here we put the brachiosaur and the diplodocus as "middle stock" because of the greater historical mediatic relevance of Apatosaurus in most nations. Sadly, Brachiosaurus recently also went through some naming troubles, but its situation isn't as severe as the Apatosaurus-Brontosaurus deal — the name Brachiosaurus remains valid, however its best-known species, B. brancai, had to be placed in a different genus, named Giraffatitan. note Giraffatitan ("titanic giraffe") was found in Africa in the Tendaguru site two decades after the U.S. brachiosaur, and is known from complete remains which show a distinctive domed skull. An impressive, 12 m tall Giraffatitan skeleton was mounted in the Berlin museum in the 1930s: this was the biggest mounted dino-skeleton until two decades ago, and the model of the popular image of the brachiosaur lasted for decades. On the other hand, the valid Brachiosaurus was long known only from fragments, and its skull was described only few years ago. This skull, which had a smaller dome than Giraffatitan's, was long classified as another kind of sauropod, Camarasaurus; ironically, it was just this skull that was put in the original "Brontosaurus" skeleton. The popular "brontosaur" is just a Mix-and-Match Critter made of Apatosaurus and Brachiosaurus.
- Entry Time: 1930s
- Trope Maker: Berlin Natural History Museum (now recognized as Giraffatitan)
The "heaviest": Argentinosaurus *Argentinosaurus was discovered Argentina in 1993. It lived in the Early Cretaceous, 95 mya, unlike the more famous sauropods, the diplodocids and brachiosaurids, which lived earlier during the Jurassic. Argentinosaurus belonged to a group of sauropods called titanosaurs, which evolved within the Titanosauriformes (which included Brachiosaurus) and replaced the earlier sauropods worldwide in the Cretaceous. Titanosaurs are based on the genus Titanosaurus, which was first described in 1877 and used as a "wastebin taxon" since then. The classification of titanosaur genera is still in debate and many (including Argentinosaurus) are based on fragmentary remains note . New finds and further cladistic research may still change the descriptions of these animals. Titanosaurs seem to have been more compact than earlier sauropods, with shorter necks and tails, solid bones, and wider frames. At least some titanosaurs had crocodile-like skin armor; in one case (Saltasaurus) this was fully developed as bony plates similar to Ankylosaurus. On the other hand, Argentinosaurus probably didn't develop an armored skin. Length and weight estimations of Argentinosaurus are necessarily speculative, but the consensus seems to put the length at 98 ft/30 m (like Diplodocus) and the weight at about 73 metric tons (about twice a Brachiosaurus). Few people know, however, that another South American titanosaur, Antarctosaurus, has left some possible remains almost the same size of the argentinosaur, which were found several decades before. Being very scant and dubious, they have been largely ignored. Other sauropods were previously estimated as even heavier than 73 tons (see the following section) but these valuations appear positively exaggerated. Such heavy land animals would haven't even able to survive, and the blue whale still remains the official record-holder of all time (only other sea creatures could have overweighed it in the past). In 2002, a Walking with Dinosaurs special (Chased by Dinosaurs) featured a herd of Argentinosaurus. In a memorable scene, Nigel Marven hurries to place weight sensors in front of the herd as it approaches, walking straight towards the camera and messing with the viewer's perspective: a very effective demonstration of the immense size of these animals. Strangely, unlike its predator Giganotosaurus This is actually a case of either Misplaced Wildlife or Anachronism Stew; the two species lived at roughly contemporary times but were found in different formations. Argentinosaurus has not received much attention in fiction since then. Maybe because, size-related impressiveness apart, the Argentinosaurus here do nothing sensational — the adults continue to walk apparently unmoved after the Giganotosaurus bring down one of their young. On the other hand, Argentinosaurus became popular among dino-fans for being described as "the biggest ever dinosaur" in the show. But this is not an isolated case. Several other sauropods have at one point been described the same way since the very first sauropod discoveries. One of the first was Atlantosaurus "Atlas lizard". Many others followed since then, with scientists seemingly competing with each other for who coined the most awesome name — Argentinosaurus is one of the rare exceptions, meaning simply "lizard from Argentina".
- Entry Time: 2002
- Trope Maker: Chased by Dinosaurs
Size matters *Let's face it — paleontologists are people too. While they carefully excavate fossils in some dusty badlands location, or sort through boxes of collected fossils in chilly museum basements, they can't help but secretly hope to be the one who discovers or describes Badassosaurus mynamii. Sometimes they do strike gold. Most of the time, they report an unremarkable animal and get the satisfaction of a job well done but very little glory. Then, there are cases like these...
- Supersaurus the "Super-lizard" was found in Colorado in the 1970s. It was described from a few bones as a brachiosaurid of unusual size, twice as long as Brachiosaurus, and hailed as the first sauropod "bigger than the brachiosaur". When more remains were found, Supersaurus was reclassified as a diplodocid, longer and more massive than those previously known but not excessively so.
- Ultrasaurus the "Ultra-lizard" is a story written across The '80s and The '90s about two sets of bones and one name. The U.S. set (a bit of backbone and a shoulder girdle, also from Colorado) was in 1979 described by James Jensen as Ultrasaurus, the largest dinosaur ever... to the press, not in a scientific paper. It was depicted as a brachiosaurid 30 m long, 16 m tall and with a weight up to 130 tons, making it even bigger than the former record-holder, Supersaurus (which, by the way, was also described by Jensen). The South Korean set (a bit of backbone and an upper forearm) was described a few years later as an Ultrasaurus. This, however, prevented the U.S. animal from being called Ultrasaurus officially, so they had to settle for "Ultrasauros" in 1991. It was still the largest dinosaur, though. Well, at least for a few years, until it was realized that the U.S. set of bones was actually from two different animals, a Supersaurus and a Brachiosaurus: the name "Ultrasauros" was consequently discarded in favor of Supersaurus. Before that the U.S. ultrasaur showed up in some documentaries, and is cited in Calvin and Hobbes as well as in the first JP novel (which mentions the following example, too).
- "Seismosaurus". In the early 1990s Ultrasaurus had to face a rival for "the biggest" title: the "Seismic lizard", popularly nicknamed the "Earth-Shaker". A New Mexican diplodocid with an estimated length of 177 ft/54 m and an estimated weight of 112 tons, which makes it almost twice as long as a blue whale, and almost two thirds of the blue whale's weight. Impressive? Well, when other experts got a look at it they determined that the size calculation had been thrown off by misplaced vertebrae, that 95-110 ft/29-33 m was a more accurate estimation, and that the seismosaur was simply an old, well-grown Diplodocus. The name Seismosaurus is now discarded.
- Sauroposeidon (the god Poseidon was also known as the "Earth Shaker", geddit?) was described in 2000 based on four extremely elongated neck vertebrae found in Oklahoma (which were, incidentally, first thought to be petrified logs). If it was a brachiosaurid, it might have had the longest neck of every creature ever (even longer than the neck of Mamenchisaurus). However, data published in 2012 puts it closer to the titanosaurs. Living in Early Cretaceous USA along with Deinonychus, some portrayals have depicted deinonychosaurs bringing down adult Sauroposeidon.
- Paralititan ("titan of the swamps") is one of the several newly-found titanosaurians cited as possible contenders of Argentinosaurus for "the biggest" title (another example is Puertasaurus). From the same habitat as Spinosaurus, it is known from a bit of backbone, a shoulder girdle, and an upper forearm. By comparing the bones with the skeleton of a more complete titanosaurid such as Saltasaurus, Paralititan appears to have been 85 ft/26 m in length and have weighed 59 metric tons. You know, unless it turns out to be a mistake.
- Bruhathkayosaurus. Found in India in the 1990s, this one is, for now, the Up to Eleven example. Described as a titanosaurian sauropod and believed to have weighed 126 metric tons, it was until recently estimated at up to 220 tons, even heavier than the blue whale. However, its formal description is extremely inadequate, and it is speculated that the leg and hip bones found are actually petrified wood. But this is not all: our exotic-named giant was initially regarded as a theropod. Imagine a carnivorous dinosaur 50 times heavier than a T. rex... Unfortunately, Bruhathkayosaurus must now join Amphicoelias below as another "one that got away"; it seems that its bones were never properly stored out of the elements and got washed away during rainstorms.
- Amphicoelias fragillimus was a diplodocid that may have been 190 ft/58 m long, and weighed 120 metric tons (making Diplodocus look like a Labrador retriever in comparison)... but the only find since 1878 is a single vertebra, which has been lost. note This discovery was largely forgotten until its original description made by Edward Cope became more widely-known in the last decade, mainly thanks to the Internet. Considering Cope's efforts to outcompete his archrival Othniel Marsh as "the greatest dino-hunter", it can't be excluded that he intentionally upsized his vertebra in its drawing. Then again, Marsh would have taken every opportunity to expose the fraud to ruin Cope's reputation, and the fact he didn't strongly suggests that the vertebra was real. There is also the distinct possibility that the fossil may not have been of a diplodocid at all, but a member of a more basal sauropod group, in which case the scaling techniques used to get its record-breaking size would be inaccurate.
- The Longest: Amphicoelias if it really existed and was a diplodocid; Diplodocus-Supersaurus-another diplodocid otherwise.
- The Tallest: Sauroposeidon if it was a brachiosaurid; Brachiosaurus-Giraffatitan-another brachiosaurid otherwise.
- The Heaviest: Bruhathkayosaurus if it really existed; Argentinosaurus-Puertasaurus-another titanosaurian otherwise.
- Entry Time: 1970s
- Trope Maker: Sensationalism in media and wishful thinking among paleontologists.
Other sauropods:Sorry, these ones aren't here. If you're looking for Camarasaurus, Mamenchisaurus, Saltasaurus, Cetiosaurus, Barosaurus, and others, see here.
Prosauropod means "before the sauropods". Living in the Triassic or Early Jurassic periods, prosauropods were among the very first dinosaurs to appear, and the first dinosaurs to reach elephant size. Some of them may have been the ancestors of the sauropods. The name Prosauropoda isn't formally used nowadays (members of the group are now referred to as basal sauropodomorphs). Plateosaurus, considered the prototypical "sauropod predecessor", is the most common in the fossil record and one of the largest as well.
The first giant: Plateosaurus *
Lived 216-199 million years ago, in the Triassic Period. Plateosaurus is one of the scientifically better-known dinosaurs, and also the most abundant dinosaur in European fossil record. More than 100 specimens are known, and even a "graveyard" in Southern Germany. Plateosaurus was also one of the first dinosaurs described, even before the word "dinosaur" was invented, but Owen didn’t include Plateosaurus in his new group (its first remains were very fragmentary). When the genus was being classified into Dinosauria, it was first placed in the theropod branch and thought carnivorous; later, was moved to the prosauropod group. Its adult size was astonishingly variable, from 16 ft/4.8 m up to 33 ft/10 m, and its weight ranged from 600 kg to 4 metric tons. At a first glance, Plateosaurus looks like a cross between a diplodocid and a theropod. The general body shape was sauropod-like, with a small head, long neck, sturdy body, and long flexible tail (and also the typical thumb-claws). The limbs and stance were theropod-like; it was bipedal, walking on hind legs that were slightly folded, rather than pillar-like. The hindfeet had distinct digits with a claw on each. The neck was shorter and more flexible than a typical sauropod neck thanks to its shorter vertebrae, recalling the necks of some theropods a bit. The head was rather theropod-shaped too, but their teeth were small and blunt, apt to grabbing vegetation instead of tearing meat. The closer relationship with sauropods is betrayed by one detail: the hands and feet of the prosauropods had five digits each like sauropods, while true theropods lost the fifth digit both in their hands and their feet (except for the most primitive controversial theropods, like Herrerasaurus and its relatives, which had five digits on their hands/feet). Science Marches On has been a strong factor within Plateosaurus portrayals. When believed a theropod it was depicted with a tripod stance like all large bipedal dinosaurs; one example could be in Fantasia. After being classified as a sauropod relative, the plateosaur has usually appeared as a slow quadruped but able to rear up its hindlegs like diplodocids, either to reach higher foliage or for defensive purpose (like in Walking with Dinosaurs). The exclusively bipedal portrait re-emerged only very recently, and today scientists believe Plateosaurus was capable of rapid runs if necessary. It may have defended itself with its thumbclaws. The plateosaur's large size could have evolved to avoid predation by the carnivorous dinosaurs (which were still small and gracile at the time); the only predators that were possibly able to defeat the adults were basal archosaurs such as the contemporaneous Teratosaurus. The two stock Triassic dinosaurs, Plateosaurus and Coelophysis, are among the most abundant in fossil record but among the least common in pop culture. Plateosaurus appearances in fiction are very rare; in documentaries, it is usually shown only to emphasize the dinosaurs' rise to power, as in the aforementioned Walking with Dinosaurs. Even though some Plateosaurus-looking dinosaurs occasionally crop up in TV (such as Dino), they are more likely humanized sauropods or Mix-and-Match Critter things.
Other sauropod predecessorsSorry, these ones aren't here. If you're looking for Massospondylus, Anchisaurus, and others, see here.
Armor-bodied plant-eatersThe Thyreophorans ("shield bearers"), were a group of dinosaurs notable for their body armor made of bony plates covered with horny sheaths. Many also developed weapons on the tips of their tails. All but the most primitive forms were massive quadrupedal animals belonging to one of these two groups: stegosaurians and ankylosaurians.
Stegosaurians were small-headed, mostly Jurassic herbivores that developed large bony plates along their backbone for uncertain purposes, and had pairs of spikes on their tail and sometimes on their hips or shoulders as well. Stegosaurus is the namesake of the group as well as one of the largest known members.
Too dumb to live?: Stegosaurus ***
Lived in Late Jurassic North America, 155 to 150 million years ago, and was discovered during the Bone Wars like several other Stock Dinosaurs (Allosaurus, Diplodocus, Apatosaurus etc.) note One of the most easily-recognized dinosaurs thanks to its bony plates, spiked tail and distinctive silhouette, Stegosaurus has always been one of the most iconic dinosaurs of all, along with T. rex, Triceratops, and a token sauropod. It is regularly portrayed both in films and in cartoons, though usually with a less important role in respect to sauropods and carnivores. The several Stegosaurus species ranged from 24 ft/7.5 m up to 30 ft/9 m long, and weighed from 1.5 up to 5 metric tons. Its plates and deep body made it looking bigger than it was when seen from the side: actually, the stegosaur’s body was laterally-flattened, and not so heavy as it seems. Its limbs were pillar-like; the front legs were much shorter than hindlegs, and the neck was set low above the ground (but not the degree seen in old portraits). Despite its overall size, Stegosaurus had a remarkably small head, with room for only 2.8 oz/80 g of brain (often stated as "walnut-sized") note . This has made it the most iconic dinosaur within the "Dinosaurs Are Dumb" subtrope (though sauropods are not far away). The small brain does not mean that stegosaurs and sauropods were witless, though. And they didn't have a secondary brain in their hip region as is often stated; the extra space there probably accommodated the nerves for the hindquarters. The back plates were the most distinctive stegosaurian feature, but it isn't entirely clear what their purpose was. It was debated whether the plates were covered in horn or in skin, but a study on stegosaur skin impression suggests the former is more likely. Defense, thermoregulation, and display (mating or threat) are the classic hypotheses, but we haven’t definitive proof for any. The early theory that they were used for armor is the most unlikely: the plates were dermic structures not attached to the skeleton, and they were irregularly placed to be used as armor and would leave the animal's sides unprotected. Although if covered in horn they might have had sharp edges, which would make them effective as defense. The "solar panel/radiator" theory was the most followed until recent years: it could explain the vessels, and also the singular arrangement of these plates — they were asymmetrically-placed, giving more surface to solar rays. This theory is still a possibility, as studies on crocodilian scutes show they have usage for thermoregulartory purposes. Walking with Dinosaurs popularized the third theory, showing a Stegosaurus reddening its plates and scaring an Allosaurus away. However, if used for display, they might also have had the function of making the animal look larger or communicating with others of its kind. Even the configuration of these plates was until recently debated. Though Stegosaurus has left dozens of specimens, they are usually found with misplaced plates, making them a sort of puzzle to rebuild. All combinations were proposed, from a single line to two paired lines. One early theory was they were flat on the back like tiles: this gave to the dinosaur the odd name Stegosaurus, "roof-lizard". The first still-articulated stegosaur skeleton was found only in the 1990s, and shows alternated plates. Stegosaurus' tail was muscular and flexible, and the animal may have been able to rest it on the ground to assume a tripod stance and reach higher vegetation (this is not certain, however). When swung from side to side, this tail made a powerful weapon against enemies. Near the tip of the tail was a group of four long spikes known as the thagomizer, a term that originates from a Far Side cartoon, later adopted by the paleontological community (you can find it used in serious scientific publications) in an even more awesome case of Ascended Fanon than "raptors". Another curiosity, though, is rarely mentioned: Stegosaurus had small scutes on its hips and tiny osteoderms under its throat. Also, a study on one Stegosaurus species, S. sulcatus, suggests that a spike found alongside a specimen may be actually from the shoulder and not from the tail as previously assumed. If this is true, then Stegosaurus had shoulder spikes like its relatives after all. It has recently been suggested that Stegosaurus was sexually dimorphic but others have cast doubts on this. If you see Stegosaurus in popular media, don't be surprised to see inaccuracies. To this day, it may be shown with paired plates or even plates in a single line, instead of zigzagging in two lines, and they may be round or triangular instead of pentagonal. And its tail may have two, three, five, six, or even eight spikes, or none at all. These spikes may be shown as much shorter than in reality note . In some cases the neck is unrealistically long, like Dinny in Alley Oop, making it resemble a cross between a stegosaurian and a sauropod. The body may be shown as very low-slung and fat (even when seen from the front), and the legs are often stubby. Stegosaurus may often be depicted with a turtle-like face instead of a horse-shaped one like in real life. Occasionally, it is shown with a beak full of teeth or even worse, no beak. In many old films, Stegosaurus is shown as a sorta "predestined loser" against big meat-eaters like Allosaurus, Ceratosaurus or Tyrannosaurus, being too slow to defend itself effectively. In modern portrayals, however, Stegosaurus more often wins fights with the aforementioned carnivores, like in Walking with Dinosaurs, as it is now considered to be agile and flexible in spite of its slow running speed and heavy body.
Porcupinosaur: Kentrosaurus *... Indeed, if you're watching a film or even a TV documentary, good luck if you’ll ever find a stegosaurian which is not Stegosaurus. However, if you do, it will probably be Kentrosaurus. Only half the length of Stegosaurus, its overall body shape was almost identical to the latter... except for the armor. The usual plates on the neck and back were much smaller and paired (not zigzaging), gradually becoming spikes on the hip and ending with at least five pairs of true spikes on the tail. But this is not all, Kentrosaurus had also a pair of isolated spikes arising from its shoulders. A Late Jurassic animal like Stegosaurus, Kentrosaurus was discovered in the 1910s in the same East African site along with with much bigger dinosaurs like Giraffatitan. Dozens of Kentrosaurus skeletons have been discovered, but with plates/spikes scattered away (as usual among stegosaurs): thus, scientists once thought Kentrosaurus side spikes were on its hips instead of its shoulders. That's why classic dino-portrayals show it with spikes protruding from the pelvis instead of from its forequarters. One mention about [mis-]spelling: Kentrosaurus should never be confused with Centrosaurus . Both names mean "pointed lizard", but the "points" of Centrosaurus were on its head: it was a ceratopsian. In some old sources Kentrosaurus is known as "Kentrurosaurus" ("pointed-tailed lizard"), but this name is now invalid.
Other stegosauriansSorry, these ones aren't here. If you're looking for Tuojiangosaurus, Dacentrurus, and others, see here.
These are the most well-armored among all the dinosaurs (sometimes called the armored dinosaurs), with low frames, quadrupedal stance, strong short legs and armor consisting of bony plates covering the upper part of their bodies. They were herbivorous and mostly lived during the Cretaceous. They aren't as common as stegosaurians in works, but still crop up semi-regularly both in fiction and in documentaries. Ankylosaurians were once divided in only two families: ankylosaurids (clubbed) and nodosaurids (club-less). Ankylosaurids had a broad head, their armor plates formed a keratin-covered shell with short spikes in many directions, and they had a tail club except for the most primitive forms; proper nodosaurids had a narrow head, rows of osteoderms on their backs and flanks, and longer spikes jutting out sideways. In recent years a third subgroup has been recognized as distinct from nodosaurids: the polacanthids, variably classified as either closer to ankylosaurids or to nodosaurids.
An (un-)well-known critter: Ankylosaurus **Lived in North America around 66-65 million years ago alongside Tyrannosaurus and Triceratops at the end of the Dinosaur Age. Discovered in 1908 in Alberta, its actual size is uncertain, extimated from about 20 ft (6 m) long up to 9-10 m, but the latter is more likely: if so, it's the biggest known ankylosaurian, possibly weighing up to 6 metric tons (as much as a male African elephant), in the same weight range as Triceratops and the biggest Stegosauruses. Despite its size, its remains are quite scant, with still no complete skeletons found, and other relatives (for example Scolosaurus) are more common as fossils. Ankylosaurus' iconic status among ankylosaurians could be explained by its sheer size and because its own dinosaurian group is called by its name. Ankylosaurus was also one of the most strongly-armored ankylosaurians — several sources have described it as a "living tank". Ankylosaurus has been famous since the 1940s as the Up to Eleven example of an armored dinosaur. In both fictional and documentary media it is often portrayed in a battle against T. rex (similarly to Triceratops). In these struggles the ankylosaur is seen defending itself by sheltering under its impenetrable bony armor, and using its tail-club like a Medieval mace, breaking the legs of its opponent and making it fall down. This might be Truth in Television, even though tyrannosaurs almost certainly didn't prey upon adult ankylosaurians frequently (hadrosaurs were much more abundants and armor-less). Despite their heavy build and short legs, they may have been able to charge the carnivore like a rhino. Like stegosaurs, ankylosaurs tend today to be portrayed as more agile and active in fights now than in the past: in Walking with Dinosaurs one easily wins the struggle (despite being shown as a very slow-walking animal), delivering to the carnivore a fatal blow with its tail-mace. When the tyrannosaur is shown winning the battle, it's seen "overturning" the ankylosaur to expone the soft vulnerable underbelly and deliver the fatal bite there. Most herbivorous or omnivorous dinosaurs are often depicted in media as gregarious animals: scenes involving herd-dwelling sauropods/ceratopsians/stegosaurs/hadrosaurs/iguanodonts/ornithomimids are a very common sight. On the other hand, adult ankylosaurians are always depicted as loners. This is realistic, though, because their fossils are more rare than those of other large herbivores and almost always found isolated. One rare exception is a group of about eight juvenile Pinacosaurus found together, which probably died at the same time during a sandstorm. Ankylosaurus probably retains the sad record of being the worst-known Stock Dinosaur. Even in documentary works, its size, shape, and composition tend to be pictured incorrectly, often with traits from other ankylosaurian species. The incompleteness of the remains only partially justify this. One common mistake is to leave out the tail club, or to have it shaped incorrectly — for example, adding spikes to it. A famous example of the latter is the "Ankylosaurus" (actually a Scolosaurus) painted by Zdenek Burian defending itself against a tyrannosaurid: it is undersized and has two spikes on the tip of its tail. When based on Real Life fossils, the club usually appears two-lobed like that of a Euoplocephalus (a close relative commonly depicted in popular dino-books), instead of elliptical. The bony covering on its back should be a snugly fitting mix of large and small plates and be interspersed with short spikes. Many classic portrayals, on the other hand, show long spikes only on the sides, similar to the related nodosaurids. Other portraits go even further, showing totally spikeless Ankylosauruses (see the aforementioned finale of Walking with Dinosaurs). Finally, the broad head should have four horns behind the eyes and the ends of the mouth. Ironically, one of the few correctly-shaped ankylosaurs in cinema is the dog-like Url from Disney's Dinosaurs (he was highly undersized, but this may be justified if he was young). Many other inaccuracies seen in ankylosaur portrayals are substantially the same as the stegosaurs (see above). Being related to each other, stegosaurs and ankylosaurs shared many features even in Real Life. They had the typical ornithischian jaws, with teeth only on the back and a toothless beak on the tip. However, their beaks and teeth were weaker than other ornithischians (ceratopsians, ornithopods); they may have chewed only soft plant material near the ground-level, and/or swallowed small stones to aid digestion, like sauropods. Even though were much smaller-sized, stegosaurians and ankylosaurians tend to be shown as slow-moving as the sauropods: ex. the aforementioned Url which has the slowest pace among all the dinosaurs of its herd (just as slow as its companion brachiosaur Baylene). Pre-Renaissance depictions used to portray ankylosaurians and other four-legged dinosaurs with splayed legs and dragging tails. Actually quadrupedal dinos had erect limbs (among them only sauropods had true claws), and footprints show they usually kept their tails above the ground when walking around. Of course, expect to see splayed-limbed ankylosaurs even in relatively recent works — Rooter of The Land Before Time appears even slower than the sauropods of the same film (the latter have correct upright limbs, though).
- Entry Time: 1940s
- Trope Maker: "The Age of Reptiles" mural
Gimme the club: Euoplocephalus *Ankylosaurus was the first clubtail recognized (1908), but, again, its remains were very scanty; however, its record size (more than 10 m long) soon made it the prototype of the ankylosaurians. But North America was home for other clubtails as well, which lived slightly earlier than the namesake of the group (though all lived in the Late Cretaceous): the traditionally most-portrayed among them has been Euoplocephalus (NOT "Eu-plocephalus" please). Euoplocephalus lived 75 million years ago in Alberta (about 8 million years earlier than Ankylosaurus), was 6-7 meters long and weighed around 2-3 tons, about the same weight of the neighboring ceratopsids (Centrosaurus, Styracosaurus, Chasmosaurus etc.), but noticeably smaller than the more recent Ankylosaurus. note . Like Ankylosaurus, it too had armor plates arranged in rows along its back; but Euoplocephalus' armor was traditionally thought to be awesomely more complex, and may even be described as "artistic". A couple of flat bumps were on the neck, and two pointed spikes protruded from the shoulders. Several other spikes were placed in regular, elegant lines along its back. Even the elbows had three small round scutes each. The head was similar to Ankylosaurus, with mosaic-like scutes on its roof, four small horns, and bony eyelids. Finally, the club was trefoil-shaped and almost resembled the club of French playing cards. A really cool animal to draw: in fact, Euoplocephalus appears as the actual stock ankylosaur in many dinosaur books. It's also worth noting that several alleged Ankylosaurus seen in books, documentaries and films tend to have some euoplocephalic traits, with conical horns instead of triangular, trefoil clubs instead of oval, and sometimes even the elbow scutes and the flat bumps on the neck. Despite this, Euoplocephalus is typically not portrayed in CGI documentaries, which will always prefer its gigantic cousin — the fact that Ankylosaurus could fight T. rex while "Euply" only had albertosaurs etc. to battle doesn’t help. Ankylosaurus and the traditionally-intended Euoplocephalus have long been considered the only Late Cretaceous North American members of the club-tailed family, ankylosaurids. Recently, two brand-new animals from New Mexico have been found; one of them has received the curious name Nodocephalosaurus (resembling a Portmanteau of Nodo[saurus] and [Euoplo]cephalus plus the usual suffix -saurus).
Club or spikes?: Scolosaurus *It seems Science Marches On can potentially hit every well-known dinosaur, suddenly making it less relevant than it used to be. Late Cretaceous North America has several example of dinosaurs that were very common in older popular dino-books but now have been "substituted" in their role by close relatives. The carnivorous Gorgosaurus was synonymized with Albertosaurus between the 1970s and the 2000s (though this has been reversed, and it is now a valid genus again); the hadrosaur Kritosaurus was revealed to be based upon the related Gryposaurus in the 1990s; the ceratopsid Monoclonius is today thought by several experts as a non-diagnostic juvenile centrosaur; and the small deinonychosaur Stenonychosaurus has been synonimized with Troodon since the 1980s. And, of course, the notorious "Trachodon" case. The couple Scolosaurus/Euoplocephalus is the latest addition in this special list. The original specimen of the latter was discovered in 1902. Between 1923 and 1929 three other genera very similar to it (Dyoplosaurus, Scolosaurus, and Anodontosaurus) were named, but these three were combined into Euoplocephalus in 1971. However, all three genera were rescued from the Invalid Box between 2007 and 2013 after showing that some patterns of armor were useful in classifying their owners. For example, the Dyoplosaurus club was different from that of Euoplocephalus, being longer than wide, while the Anodontosaurus club had pointed ends like a giant pickaxe. Dyoplosaurus was also older than most Euoplocephalus specimens, while Anodontosaurus lived after the latter but before Ankylosaurus. But it's Scolosaurus that has surely been the most relevant among these alleged euoplocephaluses. It is known from one really well-preserved skeleton from Alberta and several more incomplete specimens from Montana note . The scolosaur was about the same size as Euoplocephalus, live in the same age, had a similar head but with longer, more swept-back horns, and a club also similar in shape. The main point is: the aforementioned "artistic" armor has been found to actually pertain to Scolosaurus, while the real Euoplocephalus had less complex armor. The classic "Euoplocephalus" portrayals described above are actually based on the aforementioned well-preserved Scolosaurus found in Alberta. This skeleton, nonetheless, lacked the skull as well as the clubbed tip of its tail, making its tail look shorter and ending with a single pair of spikes (which were actually in the middle of the tail). Several old books and models have portrayed this "stegosaur-tailed ankylosaur" (wrongly showing it with much more generic armor than the Real Life fossil): interestingly, they usually named it "Scolosaurus" instead of "Euoplocephalus", but sometimes they wrongly named it "Ankylosaurus" (ex. one famous picture by Zdenek Burian which shows this critter defending itself against a Gorgosaurus). Though few noticed, even one very popular work has made the same mistake: if observed carefully, the wise "Euoplocephalus" Rooter has armor analogous to Knight's picture, and also shows the pair of spikes on the tip of its tail when he goes away, revealing it's based on a very inaccurate, old-fashioned Scolosaurus.
- Entry time: 1928
- Trope Maker: Zdenek Burian
Spiny yet clubless: Polacanthus *Some decades later, a companion was added to Hylaeosaurus: Polacanthus. English too, and also conviving with Iguanodon in the Early Cretaceous, it was also 4 m long, and also very incomplete. In older depictions, Polacanthus had very light armor, consisting only of pairs of long dorsal spikes (hence the name, "many spines"), a bony shield on its hips, and small plates on the tail. Some portrayals took it a further step and gave it a stegosaur-like thagomizer. The spiked-tailed polacanth made cameo appearances in Planet Of Dinosaurs and the film adaptation of The Land That Time Forgot as well as a more prominent role in Dinosaurs as Robbie Sinclair's friend Spike. Today we know its armor was extensive and Ankylosaurus-like (though even spikier) and with no club-like tail. The polacanth appears with this new look in Walking with Dinosaurs as a follower of Iguanodon herds.
Other ankylosauriansSorry, these aren't here. If you're looking for Pinacosaurus, Hylaeosaurus, Scelidosaurus, and others, see here.
Armor-headed plant-eatersThe most recent group of ornithischian dinosaurs, marginocephalians have been usually discovered in Late Cretaceous terrains. They were closer to ornithopods (see further) than to thyreophorans (see previous), and are divided in two very different subgroups: ceratopsians and pachycephalosaurs, unified by their armored head.
The ceratopsians were a group of dinosaurs characterized by a bony "frill" at the back of the neck. Starting as small bipedal animals like all the main dinosaur groups, they evolved towards a heavy quadrupedal body plan, while lengthening the frill and growing horns on their eyebrows and nose. Even though ceratopsians had erect limbs like every other quadrupedal dinosaur, some portrayals have shown them with splayed frontal legs. Moreover, their legs tend to be shown stockier and more "elephantine" than in Real Life. The ceratopsid family contains all the largest members of the group. Apart from the frill shape and number/length of the horns, ceratopsids shared the same basic look. They are classically divided in two subgroups: those with long frontal horns, short nasal horn and (usually) long frills; and those with short frill, no frontal horns, and (usually) a long nasal horn. Triceratops is a short-frilled member of the first subgroup, while Styracosaurus is a good example of the second one. Finally, Protoceratops has been the prototypical ancestral ceratopsian.
Mr. Three-Horn: Triceratops ***
Lived 68 to 65 million years ago in Late Cretaceous North America. It was one of the last discoveries from the Bone Wars. note Its name means "three-horned face" and is due to its most prominent feature. It was about 26-29.5 ft/7.9-9 m long and weighing about 6-12 metric tons, and was one of the biggest ornithischian dinosaurs (only some hadrosaurs were larger). Hundreds of skulls are known so far, but (oddly) not a complete skeleton. Two main species are recognized today: Triceratops horridus and Triceratops prorsus, even though up to 15 species were described at one point due to the great variability of the skull. Triceratops has traditionally been considered the largest ceratopsian; its size and abundance in the fossil record have contributed to making it the most popular one. It has always been beloved by dino-fans. Maybe because with its short tail, big head, and rhino-like body, it is the least reptilian-looking among the Stock Dinos (even in older depictions); or maybe, because of its historical reputation as "the only plant-eater able to defeat the Big Bad Tyrannosaurus rex" (even though ankylosaurs and maybe even the biggest hadrosaurs might also have been able to defeat the tyrannosaur in a fight). Its portrayal in Jurassic Park consolidated Triceratops' popularity even more: the touching scene of the sick triceratops with the caring humans around has remained in public consciousness. And how could we forget the strong temper of Cera in The Land Before Time film? Compared to other stock dinosaurs, Triceratops and its relatives have been portrayed fairly accurately. The ceratopsids in the original movie The Lost World (the Trope Maker, from year 1925) are nearly as realistic as those seen in the 1999 docu Walking with Dinosaurs (which are actually Torosaurusnote ). Thanks to their obvious resemblance to rhinos, media Triceratopses have usually been portrayed as agile and active like a modern ungulate mammal; basically, the only mistake in older depictions (other than the aforementioned issue regarding the legs) is the wide lizard-like mouth without the typical ornithischian cheeks (see also Iguanodon). Science Marches On even for "Mr. Three-Horn" however, and a third element (the completely scaly hide) has been revealed to be an inaccuracy as well. Extremely recent finds indicate that Triceratops was covered in bristles. Today, many scientists believe every dinosaur was at least partially covered in filamentous structures (just like modern mammals). Triceratops and T. rex have been shown fighting in modern works from the first dino-movies and through the whole paleoartistic tradition (including one especially iconic painting). In these battles, the percentages of victories between the tyrannosaur and the triceratops appears to about 50%. Though all this may even be Truth in Television, it's likely that the tyrannosaur preferred younger and more vulnerable prey than an adult Triceratops. Often considered the badass guy par excellence among plant-eating dinos, writers can’t resist the urge to make Triceratopses act like rhinos or even bulls. They’ll be ill-tempered, will charge everything, and may even moo like bovines. The ceratopsids’ horn structure was more like cattle’s than to a rhino’s: that is, bony protrusions covered with a horny sheath. Their function is still debated: ceratopsian horns may have simply been display devices. The frequently-seen "Triceratops goring to death a big carnivore" scene might not be realistic, and some think the frontal horns were too fragile and not pointed enough to go through flesh. On the other hand, given the keratinous sheath would have made the horn less likely to break and helped better shape it, goring may still be plausible. Another classic hypothesis is that triceratopses locked their horns like deer in head-vs-head combats, based on possible "wounds" found in ceratopsian skulls. However, only some Triceratops specimens show curved frontal horns apt for that, others had straight horns. The frill was variable, too: some individuals had tubercles on its edges, while others had smooth shields. Generally, most media Triceratopses have tubercled frills. The parrot-like jaws are rarely mentioned, to the point that some authors omit the shape from their models to make Triceratops more like a rhino or a bull. Some have gone even further, showing ceratopsians with sharp carnivorous teeth even in the tips of their jaws (especially common in some rubber toy collections). In Real Life, the ceratopsians' jaws were the strongest among all plant-eating dinosaurs, filled with sharp cutting teeth behind the parrot bill, but were arguably adapted to eat fibrous plants, not tear meat (or at least this was not the main function, see further). Some thought the powerful maxillary muscles were anchored to the frill, but this is not proven. Even less mentioned are the pair of bony knobs near the cheeks; they may have been for protecting the head furthermorely, or they may have been only for display.
Mr. Multi-Horn: Styracosaurus **
Several genera of horned dinosaurs other than Triceratops existed in Late Cretaceous North America, but only some of them have made appearances in pop culture, and Styracosaurus is the only one to do so with regularity, sometimes as substitute for Triceratops, other times together with it. Being more spectacular but less common in the fossil record than other ceratopsids, this makes it the usual Rule of Cool example. Styracosaurus lived in North America 76—75 million years ago, slightly earlier than Triceratops. It was discovered in 1913 during the second great North American "dino-rush". note About half as long as a triceratops (only 18 ft/5.5 m, weighing nearly 3 tons), the styracosaur was actually even more rhino-like. It had much longer horn above the nose but only hints of horns above its eyes. It had a round, short frill, but with several pairs of long spikes protruding from the top in a rayed manner, and shorter protuberances on the anterior edge. This sort of Horned Hairdo incidentally makes its head resemble the Statue of Liberty. No other known dinosaur had such ornamentation: other relatives had one isolated pair of spikes at the most, for example Centrosaurus. Styracosaurus had also shorter and stronger jaws than Triceratops. Some speculate styracosaurs were more sociable than triceratopses and lived in more numerous herds. Styracosaurus' frill spikes were not true horns as commonly said, but only an Up to Eleven version of those protuberances seen in almost all ceratopsid species. Even though the most common portrayal has six spikes, it seems most specimens had only four. But don’t rule out seeing styracosaurs with eight spikes or more in popular works, or even with no frill and the spikes protruding directly from the back of the neck. Why did ceratopsids have the frill? The bony core has a pair of large holes that make it less heavy but also less useful as protection (Triceratops was almost the only ceratopsid that didn’t have those holes). The frill could have been raised for threat display. Another hypothesis is that it was a thermoregulating device (like Stegosaurus' plates, Triceratops' shield seems to have been rich in blood vessels). Maybe the frill was for making the several ceratopsid species more distinctive (like the hadrosaur crests, see further). It is also possible that frills show sexual dimorphism. The styracosaur has appeared in several works since the first portrayal in 1933 (in The Son Of Kong), and is also a common feature in toys and popular books. On the other hand, recent documentaries haven't represented it so frequently. Maybe because in Real Life Styracosaurus could not battle Tyrannosaurus rex as Triceratops did, but only smaller carnivores like Daspletosaurus.
Mr. No-Horn: Protoceratops *
Protoceratops lived 83—70 million years ago in Late Cretaceous Asia, unlike the giant ceratopsids, which were mostly North American in distribution. It was around 6 ft/1.8 m in length and weighed no more than 400 lbs./180 kg. Protoceratopsids are generally smaller and more primitive than ceratopsids, and were once considered the ancestors of the latter group (hence the name, meaning "first horned face"). At first glance, Protoceratops resembled a miniaturized Triceratops — four-legged, with the same robust body, short tail, and unmistakeably ceratopsian head. However, it differed from ceratopsids mainly in having no true horns. Other differences include: a simpler frill lacking protuberances; bigger cheek-spikes; stronger parrot-jaws; and legs more adapted to running. The genders might have been sexually dimorphic (larger skulls with a nasal bump and a couple of upper "canine teeth" probably belonged to males). First discovered in Mongolia in 1922, Protoceratops is one of the most abundant Asian dinosaurs in fossil record, with hundreds of specimens discovered so far — earning it the nickname "the sheep of the Cretaceous" (given the large numbers of animals found together, they probably lived in herds). Many juveniles have also been found, and its growth pattern is one of the best understood among dinosaurs. It was the most famous Asian dinosaur until Jurassic Park made Velociraptor famous. The discoverer of Protoceratops, Roy Chapman Andrews, note attributed to it some elongated eggs which now are known to belong to Oviraptor. These were the very first dinosaurian eggs ever identified. The original crushed Oviraptor skull was found nearby (see "Oviraptor" above). A classic image in paleo-art is showing Protoceratops hatching its eggs and chasing or even trampling an egg-robbing Oviraptor. However, several nests complete with eggs were found later in Asia, which were actually laid by protoceratopses. One especially spectacular find (from 1971) consists of a Protoceratops and a Velociraptor clutched together: they were probably fighting each other when they were buried by a sudden sandstorm or a collapsing sand dune. It still remains the best evidence of a "dinosaur battle" between an herbivore and a carnivore. The Protoceratops appears to be biting the Velociraptor with its parrot-jaws, while the "raptor" is holding the protoceratops' head with its forelimbs and has one of its sickle-claws near the herbivore's throat. The real cause of the battle is uncertain, however: probably the protoceratops was defending itself and/or its offspring from the carnivore. note Despite its scientific relevance, because of its relatively modest appearence Protoceratops is less portrayed in pop-media than Triceratops and Styracosaurus. Perhaps the most well-known protoceratops is B.J., that yellow guy seen in Barney & Friends. In the much more beloved book series Dinotopia, the talking dino-character who befriends humans is also a Protoceratops.
Other ceratopsiansSorry, these aren't here. If you're looking for Torosaurus, Centrosaurus, Chasmosaurus, Pentaceratops, Psittacosaurus, and others, see here. Or here.
Unlike ceratopsians, pachycephalosaurians kept the original bipedal body plan, but evolved thick skull roofs and bony knobs on their heads for uncertain purpose. Like ceratopsians, there is the possibility that were omnivores. Needless to say, the iconic member of the group is also the biggest one, Pachycephalosaurus.
Hard-headed ram or clever friar?: Pachycephalosaurus *
Lived during the Late Cretaceous 70-65 million years ago in North America like many well-known dinosaurs. It usually shows up when an author feels like showing an "exotic" dinosaur. Its relationship with other dinosaurs has long been uncertain: originally classified as an ornithopod note , its affinity with ceratopsians was demonstrated only in the 1980s. Pachycephalosaurusnote ("thick-headed lizard") is by far the biggest known pachycephalosaur. Its actual length is uncertain: popular books often set its size at up to 30 ft/9 m; a length of 15-18 ft/4.6-5.5 m is more likely (the other relatives were not longer than 10 ft). Described in 1931 from a single skull, it was initially identified as Troodonnote , and renamed Pachycephalosaurus only in 1943. No other parts of the body have been found since then: reconstructions are typically based on a smaller, less-famous pachycephalosaurian, Stegoceras. Pachycephalosaurus is distinguished by its dome-like head which makes it look very intelligent. However, the height of the dome was almost entirely made of almost one-foot-thick bone, and its brain wasn't larger than other dinosaurs'. Its nickname "The Bone-headed Dino" is quite accurate. A number of bony knobs and blunt spikes around the base of the dome and on its nose contrasted with the smoothness of the dome to create a look of partial baldness or of a monk's tonsurenote ; hence the epithet "Friar Tuck-osaurus" in The Lost World: Jurassic Park film. Being totally bipedal, pachycephalosaurians were superficially similar to theropods, but their jaws and grinding posterior teeth were typically ornithischian (and thus plant-eating). However, Pachycephalosaurus had weaker jaws than ceratopsians or hadrosaurs and still retained small pointed teeth on the tips of its jaws which were lost in the more evolved bird-hipped dinosaurs; this would indicate the pachy had a mixed diet based on plant material with insects and small vertebrates as a supplement. Its relative Stegoceras shows small five-fingered forelimbs, a slender body, long tail, and running legs note . The body of Pachycephalosaurus probably was similar to Stegoceras, but the former being larger than the latter, its body might have had an overall stockier frame. As one of the most recent groups of herbivores/omnivores in formal dinosaur classification, pachys never appear in the oldest works. The ur-example was perhaps the 1988 film The Land Before Time, where the pachy shows up as a predatory villain trying to kill one of the protagonists with headbutts. The headbutting is a standard trait when pachycephalosaurs appear in works. Classic dino-books and documentaries from the Dinosaur Renaissance traditionally depicted males trying to impress females by ramming their heads into each other. note However, scientists found in the 2000s that the smooth domes would have slipped if struck against each other, and proposed that pachycephalosaurians bashed each others' sides and hips instead. But even this has been disputed: recent studies seem to show their necks were weaker than traditionally thought, maybe not able to withstand such an impact. Now many scientists think pachycephalosaurs simply used their dome heads to display maturity like an oversized toucan bill. Even more recently, a 2013 study found healed injuries in multiple pachycephalosaur domes, suggesting that they were used for headbutting and/or flankbutting after all. The pachycephalosaurs' real lifestyle and diet will probably remain a mystery until more complete remains will be found. The large Pachycephalosaurus was once the only bonehead portrayed in fiction. This changed in the 2000s when two smaller relatives, Stygimoloch and Dracorex hogwartsia (the latter discovered as recently as 2006) started making occasional appearances as well, thanks to their even spikier heads. A very recent theory (2009) suggests that these two horned pachys were just juvenile Pachycephalosaurus; if so, the latter will remain the only pop-cultural bonehead.
Other pachycephalosauriansSorry, these aren't here. If you're looking for Stegoceras, Homalocephale, and others, see here.
Bipedal plant-eatersThe Ornithopod group contains several ornithischian dinosaurs of different size, from Middle Jurassic to Late Cretaceous. The smallest ones were slender and completely bipedal, and probably omnivorous. The more evolved ones became bigger and returned to a partial quadrupedality, as well as becoming strict herbivores. The largest were among the most massive non-sauropod dinosaurs. "Ornithopod" means "bird-foot": they had limbs and feet similar to but (ironically) less bird-like than those of theropods. Unlike the latter, they had small mouth openings and blunt teeth for grinding plant matter. Ornithopods are the most abundant dinosaurs in fossil record; even though they lacked the thick defenses of the ornithischians mentioned above, they made up for that with either speed or sheer bulk.
Hadrosaurs are nicknamed "duck-billed dinosaurs" because of their wide, flat beaks especially evident in some species, less so in others. They all lived at the end of the Cretaceous. The biggest and most evolved ornithopods, their grinding maxillary mechanism was the most efficient of all reptiles ever, and they also developed complex prominences above their skulls with social functions. An unusually high number of hadrosaur species are portrayed in popular media, but four have received the greatest attention: Parasaurolophus and Corythosaurus thanks to their evident headgears; Edmontosaurus (called Trachodon or Anatosaurus in old media), because it's the most duck-billed duckbill, and one of the first described too; and Maiasaura, which has heavily contributed to the Dinosaur Renaissance. Breaking the usual rule, none of them is the biggest known hadrosaur.
Noisy Nature: Parasaurolophus **
Lived 76 to 73 million years ago in Late Cretaceous North America. 25 ft/8 m long and weighing 4-5 metric tons (roughly as much as an elephant), Parasaurolophus was a typical hadrosaur, with longer and stronger hindlimbs than forelimbs, three-toed feet ending in blunt nails, a long powerful tail, a small "hump" on its shoulders, a flexible neck, and the classic "duck-billed" head (although the "bill" was not as flat and wide as other relatives). Its long, backwards-pointing protrusion made its skull a bit longer than a human’s height. Even though is often called a "horn", it was actually an extension of the nasal cavities, and ended in a blunt point. note Its unique crest makes Parasaurolophus one of the most popular hadrosaurs (if not the most popular). Significantly, Parasaurolophus' remains are rarer than other duckbills. The hadrosaurs' lack of specific weapons has led to them being nicknamed "the Cretaceous antelopes". They are usually shown in dino-books and documentaries as "chosen preys" for tyrannosaurs, "raptors" and even giant crocodiles, incapable of offering resistence and obliged to flee away from them. This might be Truth in Television, but in Real Life "duckbills" were not exactly gazelle-like creatures. Adult hadrosaurs were strong and heavily-built: in a high-speed collision against a tyrannosaur, the hadrosaur had less of a chance of falling down (and would've been able to get up more easily thanks to its longer forefeet). It's easier to imagine tyrannosaurs hunted young hadrosaurs more often than adults. Like the sauropods, hadrosaurs used to be associated with water in pre-Renaissance times. The early discovery of some mummified hadrosaurs whose skin on their hands was believed to be remnants of webbing made scientists believe they were semi-aquatic creatures with literally duck-like webbed hands. We know now this skin bound the fingers together into a single, toughened "hoof" apt for walking on dry soil. Also, when on land, hadrosaurs were once shown assuming the same upright posture of an old-fashioned theropod. After the Renaissance, scientists described hadrosaurs as terrestrial animals, similar to modern ungulates but capable of shifting from a quadrupedal to a bipedal posture. Needless to say, amphibious hadrosaurs with webbed hands and upright stance still appear in recent media (see The Land Before Time). Even after it was established that hadrosaurs were mainly terrestrial, scientists still said they were more skilled swimmers than most other dinosaurs and used to flee in water to escape the (less able) giant theropods. Recent research seem to indicate hadrosaurs were not particuarly accomplished swimmers in respect to other dinosaurs, or even that tyrannosaurs were more able to move in water than the "duck-bills" — of course, this cannot be verified. Specifically regarding Parasaurolophus, countless hypotheses have been made about the function of its "horn": among them, a tool to thread its way through the dense forest foliage, or even a snorkel when swimming underwater. The latter just plain doesn't work; there aren't any holes on its tip. The most commonly-accepted scientific theory is that the complex series of tubes found within were used for amplifying calls. Scientists have even turned out to reproduce these calls, which quite resemble a brass instrument. note It's highly probable the headgear had also a display function: it might have been brightly colored to attract attention, and could have had a flap of skin stretched from it to the neck, but both hypotheses are unproved. This dinosaur has a rather strange destiny in fiction: it has appeared in almost every dino-film, but almost always in minor roles — basically with the sole purpose to increase the variety of the "dinosaur world". And don’t expect to hear its name, either. note A good example is in the Jurassic Park films. Some Parasaurolophuses are visible behind the Brachiosaurus in the famous "Welcome to Jurassic Park!" scene; some are seen in the next two sequels, too. But all these were simple cameos, and the animal is never named. note Other unnamed appearances are in Disney’s Fantasia and Dinosaurs. One rare example of a major-character Parasaurolophus is seen in The Land Before Time... at least, Ducky and her parents are officially labeled as such: they’re actually another hadrosaur, Saurolophus.
The literal duckbill has many names: the Edmontosaurus / Trachodon / Anatosaurus / Anatotitan case **
No other stock dinosaurs has had such a Mind Screw story than the Edmontosaurines (The Brontosaurus/Apatosaurus is nothing in comparison). Here, Science Marches On is to a Up to Eleven degree, coupled with a huge Taxonomic Term Confusion and I Have Many Names. Edmontosaurines roamed North America at the very end of the Cretaceous, 73-65 mya. Two genera were then recognized, Anatotitan and Edmontosaurus, but they were reunited into one in 2011. Some claim this makes Edmontosaurus the only valid name for this dinosaur, while others support the separation of Anatosaurus. By any name, these creatures included both normal-sized species (10 m) but also some of the biggest hadrosaurian species, reaching 12/13 m long — as long as T. rex and a bit heavier than it. Despite this, their size didn't preclude them to be among the rex's favorite prey. note . Edmontosaurus (if it includes Anatosaurus) is one of the most scientifically known dinosaurs. More than 10,000 known specimens (most other dinosaurs have less than 100 known specimens, but usually much fewer) show every evidence about its life, even diseases. The most striking ones are the "petrified mummies," which have preserved not only skin prints, but also hardened muscles. If you don't believe us, see here. The second find is very recent, and shows an unexpected thing: hadrosaurs had a much more massive tail than traditionally thought. If thisd be true for all dinosaurs, then many classic studies about dinosaur biomechanics should be reviewed. For example, hadrosaurs and Iguanodon are often thought mainly quadrupedal, but a heavier tail would made their center of gravity just under their hips, perfectly balancing their body on two legs. Maybe hadrosaurs mainly walked on two feet and stayed on all fours only when grazing or resting, like kangaroos. note Anatosaurus deserves the "duck-billed dinosaur" title more than any other hadrosaur, with its flat head and spatula-like beak. Edmontosaurus had a stockier head and a an undulating-edged upper bill, but was still more duckish than most relatives. Their Donald Duck-like face made these dinosaurs unusually nice-looking, making consequently ridiculous their possible portrayal in fiction as dragonlike monsters — resulting more similar to giant duck-lizards. In popular work, their "duckness" may even be strongly exaggerated, rendering its flat bill literally identical to a duck's, without any teeth or cheeks. In Real Life, hadrosaurs were not exactly toothless. Behind their bill they had up to a thousand teeth closely packed together in "batteries" and capable to grind the toughest vegetation—fossil pine needles have been found in the aforementioned mummies. Maybe they had a flap of inflatable skin on their nose to amplify their calls, but this is only a supposition. Traditionally Edmontosaurus and Anatosaurus have been considered the crest-less hadrosaurs par excellence, because their skull didn't show any bony prominence; but recently science marched on again and a specimen was discovered to have had a small, fleshy cockscomb on its head; this thing was discovered only thanks to the pietrified soft tissues found — maybe other "crestless" dinosaurs had some sort of fleshy protrusions on their head. Here’s a brief summary of the edmontosaurines’ awesome taxonomic tangle: Their first remains, isolated teeth found in USA, were named Trachodon mirabilis ("admirable rough-tooth") in 1856 — among the very first dino-remains described in North America. During the following Bone Wars, two skeletons were discovered and named Trachodon copei. Soon later, two spectacular hadrosaur "mummies" (Claosaurus annectens) were popularly referred as the "Trachodon mummies." In 1917, a gigantic hadrosaur was discovered in Alberta near Edmonton, and named Edmontosaurus regalis. In year 1942, one scientist found that Trachodon must be only used for the original teeth, and coined a brand new name, Anatosaurus ("Duck lizard"), for both the Bone Wars skeletons (Anatosaurus copei) and the mummies (Anatosaurus annectens). Before the year 1990 two well-known genera were thus recognized, Edmontosaurus and Anatosaurus, and described as two distinct hadrosaurs in dino-books and documentaries. However, in that year, new studies showed A. copei being much more different than A. annectens and E. regalis put together, and scientists changed Anatosaurus annectens in Edmontosaurus annectens. At this point the copei was the only remained Anatosaurus, but... taxonomic rules say "Anatosaurus" should indicate only the annectens. This meant it should be renamed, too. Being scientists often very nostalgic, they decided to recall it with a similar name: Anatotitan ("giant duck"). And now Anatotitan should probably be sunk into Edmontosaurus, while Anatosaurus could return valid again.Quite clear, isn’t it? Trachodon first appeared in pop-media in 1925 (The Lost World film adaptation), in which is portrayed as a prey for a giant carnivore. Since then, it became THE duckbill in public consciousness, to the point "trachodont" was also used as a popular synonym of "hadrosaur" (a bit like "brontosaur" as a synonym of sauropod). Since the "renaissance" times, Anatosaurus has become the most widely-used name. After 1990, Trachodon rapidly disappeared in pop-consciousness -– even though its ghost is still seen sometimes, like the "brontosaur" one. As it seems, the name Edmontosaurus hasn’t gone a long way in non-docu media: when an edmontosaurine appears, is simply known as "duckbill," and the crested Parasaurolophus has become the most portrayed hadrosaur today. Compensating this, edmontosaurines remain still quite common in documentary media, being the only hadrosaurs which could have met Tyrannosaurus rex in Real Life (with the possible exception of the crested Hypacrosaurus, which lived from 75 to 67 million years ago). Current dino-books usually show them with the name Edmontosaurus, while "Anatotitan" became popularized by Walking with Dinosaurs, and has also appeared in Primeval. According to the most recent researches, its proper name is either Edmontosaurus or Anatosaurus.
Greek helmet: Corythosaurus *Hadrosaurs were very diversified in Real Life. Even though they shared the same body-plan, their head was wildly diverse. They are divided in two main lineages: basically, those with hollow crests, and those without. Other than Parasaurolophus, the only hollow-crested duckbill with a significant number of appearances in pop-media is Corythosaurus. Naturally, the latter has been a much rarer sight. In theJurassic Park film series Corythosaurus joins Parasaurolophus only in the third film. Just like Parasaurolophus, good luck if you'll ever hear Corythosaurus named in fictional media. Compensating this, it has been just as common as Parasaurolophus and the Edmontosaurines in documentary works, which regularly show it with its distinctive crested look. Corythosaurus was the same size of Parasaurolophus (8 m long), and lived in Late Cretaceous North America 77-76 mya. A classic error in paleo-art is to depict these two dinosaurs living alongside Tyrannosaurus rex. Since the "rex" was discovered in more recent terrains (68-65 mya), this makes a slight Anachronism Stew case. If the artist did the research these two hadrosaurs will interact with other smaller tyrannosaurids like Albertosaurus. First discovered in 1912 by Barnum Brown note , the "cory," unlike the "para," has one of the richest records among hadrosaurs. Several complete specimens known to science, including many juveniles. The corythosaur’s cranial structure was similar to the parasaurolophus, with relatively narrow duckbill compared with "Anatosaurus" and expanded nasal bones which formed a crest. However, the Corythosaurus crest was very different than Parasaurolophus: it was laterally-flattened, round-shaped, and put upright above the head. It shape has often been compared to a Greek helmet (Corythosaurus just means "helmet lizard"), but some have (more prosaically) defined it as frisbee-like or dish-like. This crest was hollow like that of Parasaurolophus, but with less complex internal structure. It seems very different-sized and also different-shaped between genders and growth stages: adult males have the biggest, tallest and roundest ones, while those of females and youngsters were smaller and narrower, and the hatchlings were born devoid of it. Issues regarding the possible functions of the corythosaur’s crest are like those regarding Parasaurolophus. Like Corythosaurus, female Parasaurolophus could have had shorter crests than males. Even though some skulls do show some variability, Parasaurolophus fossils are too rare to make a correct comparison — maybe the different-crested specimens are just different species within the genus. Since hadrosaurian crests are so differently-shaped, experts have concluded that they had also the function to distinguish visually the different hadrosaur species/genders/growth stages from each other, just like modern antelopes with their distinctive horns (let’s face it, comparisons with antelopes do work very well when talking about hadrosaurs). Moreover, the different-sized crests made differently-pitched sounds. As trombones emit lower notes than trumpets, adult males’ voices were lower than females, which in turn were lower than youngs. Then, as French horns and bassoons have a different timbre, so would have been for Corythosaurus and Parasaurolophus.
A good mother: Maiasaura *This hadrosaur deserves a special mention. The same size of Parasaurolophus and Corythosaurus and contemporary to them, 74 mya, Maiasaura had no such striking headgear (only a small relief above the eyes), nor did it have such a wide bill like the edmontosaurines. Nonetheless, it has been one of the most important dino-finds ever. Hundreds of Maiasauras were discovered together in Montana in year 1980 by famous paleontologist Jack Horner note , in what is known today as the "Egg Mountain." His mountain showed not only adults, but also many fossilized, 6 ft wide, crater-like nests made of earth and full of hadrosaurian eggs, hatchlings of all ages, and even skeletons of embryos still inside the eggshells! Before the 1980s only few dinosaurian eggs were known to science, and parental caring among dinosaurs was still a very speculative issue. Horner's discovery was a true snapshot of daily dino-life. He noted that the youngest specimens still had incomplete limb-bones: this meant they were incapable of leaving their nests. And yet, their teeth were noticeably worn, as they were already eating tough vegetation. How could they feed on themselves? Here is the proof of parental care: only adult maiasaurs could feed the young to make them surviving until they grew larger and finally could leave their nest alone. Horner gave a Meaningful Name to his caring dinosaur: Maiasaura means "good mother lizard" (note the unusual feminine suffix -saura). It was just his deep study about this dinosaur that has given to Horner his current prestige in the scientific community. Horner and then other scientists made this possible reconstruction of Maiasaura lifestyle. Huge herds of possibly 10,000 individuals used to migrate across Western North America from the northern Canada south to Montana to winter in their island. Here, they mated, built their nests, laid their eggs, and filled their nests with decaying vegetation to keep the precious eggs warm. note After the hatching, adults feed their helpless babies with good food, moved by their cute appearance (the babies’ skulls show large eyes and short muzzles like modern mammal cubs). After having developed their skeleton, the youngsters started to search their food on their own; finally, the whole herd undertook again their migration toward the North, to pass here the Polar summer. In short, an overall behaviour very similar to many modern migrating birds. This reconstruction made the top of the Dinosaur Renaissance, definitively debunking the old "big, stupid, unfeeling, oafs" thing, and making Maiasaura just as common in popular books as Parasaurolophus Corythosaurus & "Anatosaurus" since then. Some years after 1980, the discover became known among pop-writers, too. Only... Maiasauras inconspicuous appearance was not interesting enough. Even though the "good mother dinosaur" and the whole argument are widely mentioned in the 1st Jurassic Park novel, note this was totally overlooked in Steven Spielberg's following film. Other Hollywoodians resolved the problem in another way: giving Maiasaura's behaviour to other relatives. In The Land Before Time, the hadrosaurs (actually, every herbivorous dinosaur) migrate through the lands and hatch their young in crater-like nests made of earth. This was copied later by Disney's Dinosaurs (this time the duckbills were substituted by Iguanodon). All OK? Obviously, not. We have no proof if other dinosaurs really behaved the same. It's like saying that if sparrows build cup-like nests, then every other bird must build cup-like nests just because is a bird. Mind this: have you ever seen an ostrich or a penguin brooding their eggs in a cup-like nest built on a branch? In the New Tens, Maiasaura makes its first notable film appearance in the Japanese animated movie You Are Umasou where it's shown to live up to the "Good Mother" in its name, taking in an orphaned Tyrannosaurus and lovingly raising it as its own along with its biological children.
Other hadrosaursSorry, these ones aren't here. If you're looking for Hadrosaurus, Kritosaurus, Saurolophus, Lambeosaurus, Tsintaosaurus, and others, see here.
There were many non-hadrosaur ornithopods as well, but only one of them has made significant appearances in fiction, Iguanodon (again, this is the biggest of the ensemble). If you are lucky the much smaller Hypsilophodon may also show up, but it's almost never named.
A veteran: Iguanodon **
Living 126 to 125 million years ago in Early Cretaceous Europe, this is the most iconic non-avian dinosaur from the "old continent." It’s also one of the most scientifically well-known dinosaurs, and one of the most abundant in fossil record. Iguanodon has had a special role within the stock dino-ensemble. Along with Megalosaurus, it’s the only dinosaur that has covered the whole history of scientific and popular portraits, but unlike the megalosaur, has managed to preserve its fame still today. Although nearly as big as Tyrannosaurus (10 m or more, and up to 4 tons), Iguanodon has not an expecially striking look among stock dinosaurs. Being an earlier relative of hadrosaurs (and possibly their ancestor), its shape resembled one of the latter, with three-toed hindfeet, short but strong forelimbs, long tail stiffened by bony tendons, massive body, hindlimbs much longer than forelimbs. Non-hadrosaurian traits include: the backbone not curved at the shoulder level; grinding teeth much less numerous and put in one single row on each half-jaw; a totally crest-less head; a deep, narrow beak very unlike the duck-billed one. The hands of Iguanodon contain all the "oddities" in its skeleton. The most known is the spike on its hand made of the first digit's phalanxes fused together and encapsulated in a horny sheath, usually shown in books as a weapon against enemies. And it had an opposable "pinkie" finger, maybe to grasp vegetation. Like hadrosaurs, the three central digits were fused together in a hoof-like structure and supported the weight of the dinosaur when on four legs (though incorrect freely-fingered iguanodonts-hadrosaurs often appear in portraits). We don’t know for sure if iguanodonts (and hadrosaurs) were mainly tree-browsers or ground-grazers. However, classic portaits usually show iguanodonts in the usual "tripodal" stance and browsing like a giraffe. Several paleo-works have also added a long extendable giraffe-like tongue to reach tree-foliage, but this is unlikely. Iguanodon is one of the three animals along with Megalosaurus and the ankylosaurian Hylaeosaurus which were called "dinosaurs" for the first time in history (1842), by the English paleontologist Richard Owen. Iguanodon was already identified in 1825, just one year after Megalosaurus, by English doctor and fossil-collector Gideon Mantell (it's the second non-bird dinosaur to have received a official name). It was initially described from its iguana-like teeth and few other incomplete remains: hence its name meaning iguana's tooth. But then, in 1877 about 40 Iguanodon skeletons were discovered within a coalmine in Belgium near the town of Bernissart, the very first "dino graveyard" ever found. Many other remains were later assigned to Iguanodon (often found outside Europe), but many have recently split in other genera (see Prehistoric Life). Most dinosaurs have changed their look at least once: Iguanodon has done this twice. The first attempt of reconstrution showed a huge dragon-like quadruped, and one of its thumbspikes was inaccurately put on its nose—this is justified by the very fragmentary nature of its original remains. note After the discover of the complete skeletons from the "dinosaur mine" in the 1870s, the iguanodon became bipedal and upright, but still reptile-looking, often shown with iguana-spikes running along its back, and with an overall theropod appearance. Finally, studies started in the 1970 and led by English paleontologist David Norman made Iguanodon returning quadrupedal again (though still capable to stay and run on two legs), and with cheeks hiddening the teeth in the living animal. note An excellent example of this new portrait is seen in Disney's Dinosaur, which made Iguanodon the main character in the story — exaggerating its horse-like look with fleshy lips instead of the proper bill. Even though has been extremely common in dino-books and other non-fictional media, Iguanodon has not made significative apparitions in cinema or TV before Disney's Dinosaur and Walking with Dinosaurs were broadcast during the 20th-21st century change. Rule of Cool easily explains why: with its generic look and weak weapons, it don’t bear the comparison with Tyrannosaurus rex jaws, Triceratops horns, Stegosaurus plates, "raptor" claws, or the immense size of sauropods — and some portraits could even leave the beak or the thumbspikes, making it even more generic. However, its historical and scientifical importance won't ever be deleted in dino-fans' consciousness, as no other dinosaur has run the whole two centuries of popular portraits: from Crystal Palace rhinos, to giant two-legged iguanas, up to Disneyan horses.
- Entry Time: 1852
- Trope Maker: Crystal Palace Park