Creators said as much about Merrin Dungey's casting as Ursula from The Little Mermaid, as she does not resemble the character much. But they were inspired to cast her because of her performance in Alias.
Likewise Joanna Garcia is a full eighteen years older than Disney's Ariel, being 33 when she first played her. But it's hard to imagine an actress who can capture Ariel's optimism, clumsiness and quirky attitude better than she can.
According to Rebecca Mader, she expected to do an American accent to play Zelena - since the Wicked Witch comes from an American fairy tale. But the creators insisted on using her natural accent (although this may be an example of another trope), and she was the first choice for the role. (Conversely, Bo Beep (who appears in season 4's "White Out") has an English accent, but was played by American actress Robin Weigert.
Season 2 featured several episodes in which lead actors are absent as the storyline focuses on one specific set of actors for an episode. Ruby/Red is probably the most noteworthy example, to the point where the character was quietly written out towards the end of the season after the actress was cast in another series. However, she has guest-starred in a couple of episodes during the second half of the third season. In Season 5, her departure is shown as well as what's she's been doing since then.
A bizarre case in the second half of season 3. Michael Raymond James's character has been killed off, and yet his name is still in the opening credits three episodes later. He finally reappears in "A Curious Thing" as the one who sent Hook the message to find Emma in New York by briefly leaving Rumplestiltskin's body.
Christie Laing, who plays Marian, didn't know that Marian was actually Zelena in disguise in season 4 until "Heart of Gold".
Eion Bailey auditioned for August pretty much knowing only what was in the script for his first appearance. He joked that the creators went to great lengths to hide the twist that he was really Pinocchio for as long as they could from him.
Otherwise averted with David Anders, who only signed on when he was told his character was really Dr Frankenstein. Likewise Meghan Ory was told that Red would turn out to be the wolf a few episodes before it actually happened.
Colin O Donoghue had no idea Hook was a second Dark One for the first half of Season 5. Quite justified in this case as Hook didn't know either.
According to Robert Carlyle in a DVD commentary, he and Robbie Kay were the only actors told that Peter Pan was Rumple's father until the episode when it was revealed.
David makes a comment to Snow about falling through a portal to Asgard. Josh Dallas was the original Fandral in Thor.
Eion Bailey previously played a writer in Band of Brothers. His character in that went on to write a book about sharks in real life and ended up lost at sea - which is interesting, given that Pinocchio was nearly eaten by a whale.
Actor-Inspired Element: Elizabeth Mitchell suggested that Ingrid be barefoot, as she found it hard to walk in the woods while wearing high heels.
Ascended Fanon: Due to how little the final film bore to the source material, many fans of Frozen had theorized that Andersen's Snow Queen existed in the Frozen-verse as a separate entity than Elsa, another fan theory being that Elsa's powers were tied back to the former. Once appears to be going with an idea similar to this, with Elizabeth Mitchell portraying the Snow Queen as a separate character from Georgina Haig's Elsa. Both theories were ultimately used, with the Snow Queen turning out to be Elsa's aunt.
As noted above, David Anders signed on when he was told he was actually playing Dr Frankenstein.
Rebecca Mader was a huge fan of the show and claims she screamed in public when she was offered a role. She agreed before she even knew which role it was.
Merrin Dungey likewise immediately agreed to move to Vancouver to film and dye her hair blonde, as she had been a big fan of The Little Mermaid - and jumped at the chance to play Ursula.
Georgina Haig had a reaction like this to being cast as Elsa. She describes walking around in the costume as "like being Santa Claus".
Billing Displacement: Raphael Sbarge (Archie) during season 1 and Michael Socha (Will Scarlet) during season 4 are in the credit opening as main characters despite being actually secondary. Eoin Bailey also qualifies as although his role in season 1 did become crucial in later episodes, he was only in half of them and ultimately functions more as a supporting character; like Sbarge, he was billed as recurring/guest thereafter.
Robert Carlyle hated that Neal was killed off, feeling Rumple's character lost his main motivations afterwards.
Amy Manson not-so-subtly retweeted a couple of fan tweets complaining about the Season 5 winter finale where all her scenes were cut.
Elliot Knight also retweeted fan tweets that expressed disappointment in his character being so abruptly and unceremoniously killed off.
Ginnifer Goodwin has also subtly hinted on more than one occasion that she's not happy with the direction Snow's character went.
Jennifer Morrison has publicly disagreed with the producers' character interpretations on several occasions, most notably saying that Emma taking the darkness into herself was about saving the whole town, rather than just Regina.
Michael Socha has been quite vocal that he didn't enjoy his time on the show because even though he was contracted for a full season he really didn't do anything while he was on set.
Sean Maguire also expressed his frustration with the lack of closure following Robin Hood being unceremoniously killed off. However, after talks with the creators, Sean felt content enough to return to the show and wrap things up.
Dawson Casting: Common where the younger versions of characters in the flashback sequences are played by the same actor, but there are also some completely straight examples.
Especially notable in "Tallahassee" which features Jennifer Morrison and Michael Raymond James as the teenaged versions of Emma and Neal. By the time Season 3 came around, Jennifer Morrison was too old to convincingly play Emma as a teen, and so Abby Ross was cast to make this slightly better.
Elsa is 23 years old and is portrayed by 29 year old Georgina Haig. This is just slightly less egregious than Frozen, where a 21 year old Elsa is voiced by 42 year old Idina Menzel.
Whilst played straight in Frozen (where a then-18 years old Anna is voiced by a 33 year old Kristen Bell), OUAT makes it a minor trope, since Anna is 20 years old and is played by a 22 year old Elizabeth Lail.
Nicole Munoz was twenty when she played Lily at 16. And since the show is a little vague on how much time has passed since the first episode, the adult Lily is this too. There's a seven year age gap between Agnes Bruckner and Jennifer Morrison, despite their characters supposedly being the same age.
Ruby is implied to be late teens or early twenties. Meghan Ory was nearly thirty when she first played her.
Peter Pan's age is vague, but he's meant to be an eternal little boy. Robbie Kay was eighteen.
Cora's younger self is played by Rose McGowan, who had just turned forty. Her age isn't stated in the flashbacks, but is implied to be very early twenties at the latest.
Defictionalization: The exterior shots of Storybrooke's main street are filmed on a real road in British Columbia. The signs on certain shops are covered with a Storybrooke equivalent during filming and removed afterwards. However, some shop owners let the production company keep the signs up permanently partially due to this trope and partially to make less work for the film crew (to erect and tear down the signs every week). The producers also leased two units just for the storefronts. These are kept the same year round, since they were never real shops in the first place.
Deleted Scene: The Season 5 winter finale featured a scene wrapping up the loose ends of the Camelot storyline, but it was ultimately cut from the episode.
Development Gag: In "Snow Falls", Mary Margaret tells Emma "I'm a teacher, not a nun." In the original pilot script, Mary Margaret was actually a nun.
Dueling Shows: Was initially this with Grimm, an NBC show that debuted around the same time that also focuses on fairytale characters existing in the real world. It soon became clear both shows were going in different directions. Grimm focuses on more obscure characters and creatures from the many, many lesser known Grimm fairytales, as opposed to the more famous and iconic ones in Once.
Merida's parents are both played by Irish actors. The other people from Dun Broch are likewise played by local Canadian actors. Averted only with Merida - played by Scottish actress Amy Manson - and with Macintosh - played by Scottish actor Paul Telfer.
English actor Stephen Lord puts on a Scottish accent to play Rumple's father Malcolm.
The Queen's fans are called Evil Regals, Snow White's are the Fairest, the Seven Dwarves' are Team 7, Prince Charming's are Charmers, Ruby's are Little Reds, and Belle's are Beauties. Emma's fans were originally called "Team Emma" though "Ugly Ducklings" is becoming more popular.
Snow White becomes pregnant midway through Season 3 to accommodate Ginnifer Goodwin's real life pregnancy. But in the episodes before this is revealed, you can see the familiar tricks of her wearing big coats and loose tops. The Season 3 finale, after she's given birth, features time travel to the events of a Season 1 episode. It's obvious when they switch from archive footage to newly shot stuff, as Snow gains a double chin. Luckily she was already wearing a long black cloak in the episode.
Emilie de Ravin's pregnancy was also written into Season 5. But flashbacks in Belle's episodes shoot her from the chest up. She also willingly undergoes a sleeping curse to allow Emilie to hide the bump and allow for a Time Skip.
Ginnifer Goodwin was also pregnant a second time in Season 5. They kept Snow in big coats and wrote her out of the storyline temporarily to hide it.
Mr. Gold perfectly remembers being Rumplestiltskin. Most fans probably guessed it from a mile away, though The Reveal was no less satisfying.
Surprise (or not)! Cora is the Queen of Hearts after all.
Several people guessed as early as the first season that Henry's father (Neal) is Baelfire.
Fan boards had joked since Frozen hit theatres about how it would translate to that universe. Guess who just showed up...
Many had theorized that Elsa wasn't the actual Big Bad of Season 4 and that the Snow Queen was the real villain. They were right.
A smaller case. When Merrin Dungey's Ursula appeared in Season 4 (and was referred to as "the sea witch"), a few assumed that she wasn't the sea goddess that Eric's kingdom worshipped. "Poor Unfortunate Soul" confirmed that she was just named after the goddess - though it's not confirmed whether that was her at the end of "Ariel" or the true goddess.
Lily being the daughter of Maleficent was so obviously guessed by everyone it would have been a bigger twist if it hadn't turned out that way.
Henry's middle name being Daniel was a popular theory.
A smaller one brought on by The Reveal that Zelena was really Marian in disguise. Two episodes later revealed she was pregnant with Robin's baby, which had been tossed around by fans.
After the Black Fairy showed up in Season 6, it did not take fans long to suggest that she would kidnap Belle and Rumple's son.
The Evil Queen is — well, evil. Lana Parrilla, on the other hand, is an incredibly nice, warm, lovely human being who adores her fans and fellow cast members, is a second mom to Jared Gilmore (Henry), and has embraced her "Evil Regals" with open arms.
Rumplestiltskin is... not nice. Robert Carlyle, on the other hand, is by universal acclaim the nicest person on the set. Given the temperaments of most of the rest of the cast, that's saying something.
Hook was not a very nice person initially. Colin O'Donoghue, on the other hand, is incredibly polite, kind, and is completely Adorkable.
Zelena is, of course, wicked. Rebecca Mader is a kind, geeky, Adorkable woman who is always ready to answer her fans' questions on her Twitter, and was a big fan of the show before even being in it.
Peter Pan is one of the most horrible characters on the show. Robbie Kay is a nice guy and very easygoing. When talking with fans, especially younger ones, he really goes out of his way to discourage them from comparing him as an actor to the character Pan. He makes it clear he's just playing a role and he's nothing like Pan, but he's so skilled as an actor that Pan has had a great impact on the show. You can tell these questions affect him and he doesn't want anyone looking up to Pan or comparing him to Pan, who is a genuinely evil character.
Cora is the reason Regina is evil and makes her look nice at times. When Barbara Hershey had a scene in "Sisters" that required her to shout at the actress playing Young Regina she hugged the girl afterwards.
It seems to be intentional. In Season 1, the Queen of Hearts is played by Jennifer Koenig. In Season 2, she's played by Barbara Hershey when she's revealed to be Cora. It should be noted that even when she was played by Jennifer Koenig, Barbara Hershey supplied the voice, which was hard to make out due to her speaking only in whispers.
In his first appearance, "Skin Deep" in Season 1, Gaston is played by Sage Brocklebank. In his second, "Her Handsome Hero" in Season 5, he's played by Wes Brown, and the "previously on" flashbacks have Brown inserted into the scenes from "Skin Deep".
In season 3, Sean Maguire replaces Tom Ellis in the role of Robin Hood. It's later justified when a flashback taking place before his first appearance shows Robin stealing a glamour charm from Zelena and using it to hide from Rumple, whom he double-crossed. He says that the charm will make him appear to be someone else in the eyes of his enemies, so Rumple doesn't recognize him when they meet next.
In season 4, Merrin Dungey replaces Yvette Nicole Brown in the role of Ursula. Subverted when the episode Poor Unfortunate Soul reveals that they're separate characters: Dungey's character was named for Brown's.
Rose McIver usually plays women with tragic backstories, or who have gone through Break the Cutie in some way. Tinkerbell is similar but recovers from her damage and reverts to The Cutie pretty quickly.
Frances O'Connor, who plays Belle's mother, is only fifteen years older than Emilie de Ravin.
And last, but far from least, Lana Parrilla is only one year older than Ginnifer Goodwin. This is fine in the first few episodes, since Parrilla is only known as the stepmother. But Regina's backstory episode has Parrilla play the young Regina, with a child actress playing Snow at eleven. This means there must be at least an eight year difference between the two characters.
Brave's Queen Elinor is played by an actress that looks a bit young. Granted she would have been fine for the events of the film. But here Merida and her brothers are at least ten years older.
As has been widely publicised, Ginnifer Goodwin and Jennifer Morrison have frequently been mistaken for each other so many times. So they're a perfect fit to play mother and daughter. Jennifer Morrison also shares some features with her on-screen father Josh Dallas.
Abby Ross, who plays Emma as a teen, has a strong resemblance to Jennifer Morrison too. As does the uncredited young actress who plays Emma's child self in "The Dark Swan".
There's also a very good resemblance between Lana Parrilla and Barbara Hershey, who play mother and daughter. Rose McGowan was also praised for looking uncannily like Hershey's younger self when she played the young Cora.
The resemblance between Ginnifer Goodwin and Bailee Madison is also uncanny.
During the Ruby/Granny fight in "Red Handed" August can be seen barely containing himself from laughing. Eion Bailey really was trying not to crack, but it was so funny the scene was added.
Rumple's Important Haircut in Season 6 was because Robert Carlyle had cut his hair short for Trainspotting 2.
Unreliable Expositor: Emma claims to have been 18 when she had Henry, but he was already 10 when she turned 28, meaning she had to have been 17 when she gave birth to him. In "Manhattan" (which at the most is 6 months after the pilot) Henry is 11, making Emma 16 when she was pregnant.
A common occurrence through DVR ratings. Example: the first half of Season 3's Neverland arc (episodes 1-6) was a ratings success on all fronts, but the second half not so much. A rerun of its final episode, however, brought its ratings score up to standard.
Similarly, even though the live ratings have plummeted recently, it has become one of the top shows that people rent and binge-watch on Netflix.
Apparently they wanted Lady Gaga as the Blue Fairy.
The Sheriff was meant to be Sherlock Holmes, and his curse in Storybrooke was that there was no mystery to solve. The plan fell through due to an issue with acquiring the rights. Not to mention the fact that there are already twoSherlocks on TV at that point in time.
Hook was supposed to have a larger role in the second half of season two, but it had to undergo a major rewrite due to Colin O'Donoghue breaking his leg. The writers had to cut almost all of Hook's scenes.
Regina and Henry's relationship from season 2 onward is only a result of Lana Parrilla (Regina) pushing the writers for it. The same is true for Regina and Robin's relationship, somewhat...she only pushed for a romance for Regina, the choice of Robin Hood for the love interest is the writers' doing.
Belle was originally intended for only one episode, but she was brought back by popular demand. Similarly, Anton the Giant was originally going to only be in his nameless debut, "Tallahassee", but the staff liked him enough to bring him back and expand upon him in "Tiny" and "Lacey". Another similar case is Blackbeard, whose death in his debut, "The Jolly Roger", was later retconned because the writers really enjoyed what Charles Mesure brought to the role and thus wanted to use the character in more stories.
Captain Hook was originally supposed to appear in season 1, but the writers were unable to acquire the rights to Neverland in time.
Dracula was considered as a character for season 1, but they went with Dr. Frankenstein instead.
Ruby had more plots planned but the ever-increasing character load in season 2 left them no way to feature her. Season 5 re-introduces her.
Amy Manson claims she was offered to make Merida a series regular. She declined, not wanting to move to Canada to work exclusively there.
Sean Maguire stated that if Season 5 had been the final season and the show not been renewed, then Robin Hood wouldn't have been killed off, since that was done to kick off a plot for Season 6.