Characterisation Marches On: The first book despict her as a soldier of apparently average intelligence compared to 56. Later books emphasize her curiosity and despict her as incredibly smart for an ant, capable of understanding human culture and even adapt some of their concept to Ant society. By the time of Book 3, she had been so influenced by human culture that many Ants consider her a Cloud Cuckoo Lander.
Did You Just Punch Out Cthulhu?: From a Humans Are Cthulhu perspective, she is the only known Ant to have defeated humans. Though her success in Book 2 was mostly thanks to luck (the human happened to be allergic to bees), the second Book has her defeat Maximilien Linard.
Easy Sex Change: more like easy sex add. All she had to do was eating some wasp meal and she became a fertile female.
Genius Bruiser: From the Ants' perspective; she is despicted as big and strong by Ants' standards, but it's also pretty clear she is one of the smartest in her colony (at least in the two later books; she was more average before).
The Cassandra: The ants don't want to listen to him when he tries to warn them about the ant-slaughter he has witnessed, since he fails to bring proof. The ant Mibs didn't help.
Decoy Protagonist: he is the first ant character introduced in the book, and the one followed for most of the beginning of the story, making you think he might be the hero... until he is killed right toward the middle of the book.
Took a Level in Jerkass: She grows more and more irrational about her faith in Humans as Gods over the course of Book 2 and 3. By the third book, 103 is forced to have her kicked out of the colony because of this.
Easy Sex Change: Similarly to 103, gets turned into a male in the third book.
Out with a Bang: 103 knowns that, since male ants can't survive mating, she can't allows herself to have sex with 23 in the third Book, since she cares too much about him. They eventually do it when it appears they are probably going to die anyway.
Irrational Hatred: Her hates of humans reach such level that she eventually turns against 103 and tries to kill her just for possibly be in league with humans. All of this while they are in the middle of a human city, with her, 103 and 23 as the only survivors of the crusade.
Knight Templar: She is even more hateful toward Humans than Chli-Pou-Ni is.
Serial Killer: Only in the second book, and he wasn't confortable with it.
Well-Intentioned Extremist: He tried to prevent the creation of an insecticide intended to exterminate all insect life (especially ants) by gruesomely killing all the scientists working on it with an army of ant robots.